Wheeler and Wheeler — Myrmicine Larvae 145 
other Myrmicinae, they do not spin cocoons but form naked 
pupae, they nevertheless possess huge salivary glands. Even 
in the very young larva the salivary receptacle on each side 
is full of a clear liquid secreted by the large cells of the two 
branches of the gland. In the nearly full-grown female 
larva the glands are very voluminous and their lumen and 
that of the receptacle full of secretion shown as dark, com- 
pact masses in the figure, which was, of course, drawn from 
a specimen hardened and dehydrated in alcohol. As such 
an amount of saliva would hardly be necessary for digestive 
purposes and as it is not used in the form of silk by the 
full-grown larva, it probably serves as a store of food for 
the nurses. The Paedalgm larvae, therefore, would seem 
to resemble the repletes of honey ants . . . except that the 
food for the workers is metabolized and stored as saliva by 
the larva, instead of merely being ingurgitated and stored 
in the ingluvies, or crop by a certain number of workers. 
From the fact that other Myrmicine ants, although they 
spin no cocoons, often have well-developed salivary glands, 
we may infer that these organs have much the same func- 
tion as in Paedalgus” (pp. 301-302). Fig. 5 on p. 303: A, 
very young larva in side view showing internal anatomy; 
B, nearly mature female larva in side view showing salivary 
glands. 
Wheeler, 1922: “They are white, nearly spherical, with 
short neck, small head, and very feebly developed mouth- 
parts, indicating that they are fed by the tiny workers with 
regurgitated liquid food. They are . . . covered uniformly 
with short, stiff, sparse hairs, each of which has two re- 
curved branches. Even in alcohol, the larvae cling com- 
pactly together in masses by means of these hooks. When 
stained and cleared, the larvae are seen to possess unusually 
voluminous salivary glands. The youngest individuals, 
scarcely 0.2 mm. long, have the receptacle full of clear 
secretion. In older larvae, the secretion after dehydration 
forms great masses in the receptacles and lumen of the 
glands. As these organs are not used in spinning a cocoon, 
it is very probable that the secretion ... is elaborated and 
used as a food for the workers (trophallaxis) ” (pp. 179- 
180). Fig. 43 on p. 179 = Wheeler, 1918, Fig. 5. 
In this same article Wheeler concluded (pp. 118-119) 
