82 
Philippine Journal of Science 
1920 
fig. 13, Plate 2. This might be interpreted as evidence of a type 
of precocious nuclear division along the lines seen in certain 
of the hypotrichous ciliates. It is hard, however, to reconcile 
this view with the absence of evidence of division of the ble- 
pharoplast, parabasal, and flagellum which should occur before 
nuclear division. Some of the British protozoologists, who are 
quite familiar with the trypanosomes, hold that this appear- 
ance, which they have seen in other forms, has nothing to do 
with fission. 
Other species than those already mentioned have been found 
to possess posteriorly situated nuclei, among them being Tryp- 
anosoma equiperdum, T. equi, and T. pecaudi. 
Hartmann and N611er(2) in their recent paper on the cytology 
of Trypanosoma theileri figure (Taf. 14, fig. 1) a trypanosome 
with the parabasal immediately in front of the nucleus which, 
however, is situated at the middle of the animal. It was a cul- 
tural form, and they report no such appearance in fresh blood. 
Wrublewski,(iO) so far as we have knowledge, comes nearest 
to the conditions obtaining in our case. In his description of 
Trypanosoma wrublewskii Vladimiroif and Yakimoif, which is 
found in the blood of the Lithuanian bison, Wrublewski men- 
tions the finding of trypanosomes, the trophonuclei of which lay 
in the mid-portion of the body which was broadened at that 
point, the kinetonucleus (parabasal) lying in front of the tro- 
phonucleus in each case. As the trypanosomes Wrublewski 
studied were found in blood taken from the hosts after death, 
his conditions will be seen to correspond somewhat with ours. 
Martini (6) describes a trypanosome recovered from the blood 
of cattle in the Philippine Islands which may be identical with 
the organism discovered by us. Unfortunately, however, he 
fails to give measurements of the parasite and it is, therefore, 
impossible to form any definite opinion on this point. Forms 
he figures showing the parabasal lying anterior to the nucleus 
were found in blood cultures, and he says nothing concerning 
their discovery in the blood of the host during the life of the 
latter. 
Whatever the interpretation of these forms, we are not in- 
clined to regard them as crithidial forms in the true sense of 
the word. They appear to develop by posterior migration of the 
nucleus rather than by anterior migration of the parabasal with 
accompanying attenuation of the anterior end of the trypano- 
some, characteristic of the assumption of the crithidial stage by 
a trypanosome. It would be futile at this time to speculate at 
