16, 8 
Taylor: Philippine Amphibia 
233 
tigerina Daudin and R. macrodon Kuhl. Stejneger recognized 
that the large-toothed species of the Philippines is distinct from 
R. macrodon and named it R. magna. Rana vittigera and R. 
moodiei have long been confused with R. tigerina, but the char- 
acters which obtain in the first two groups show clearly that 
they are very distinct from R. tigerina. 
It was with great hesitancy that I united the two species, 
Rana varians Boulenger and R. sangumea Boettger. I have not 
examined the types, but a comparison of the type descriptions 
shows no differences warranting separation. My opinion was 
confirmed by a comparison of specimens from the Calamianes 
and Palawan. 
Key to the Philippine species of Rana Linnseus. 
a\ No gland on upper arm. 
P. Fingers and toes not or but slightly dilated at tips; no bony teeth 
in lower jaw. 
c\ Eyelid* much wider than interorbital distance; no distinct canthus 
rostralis (large frog). 
cf. A flap of skin on outer side of fifth toe and metatarsal. 
R. moodiei sp. nov. 
tf. No skin flap on fifth toe and metatarsal.... R. vittigera Wiegmann. 
c^ Eyelid about equal to interorbital space. 
cT. Toes almost completely webbed; skin granular (small frog). 
R. mindanensis Girard. 
(f. Toes about two-thirds webbed; a narrow dorsolateral fold. 
R. palavanensis Boulenger. 
c’. Eyelid much narrower than interorbital space; skin smooth; toes 
one-half webbed (small frog) R. parva sp. nov. 
6’. Fingers and toes distinctly dilated into small disks, without a groove 
around the edge; two well-developed bony teeth in the anterior 
part of lower jaw. Canthus rostralis distinct, angular, 
cb Toes entirely webbed; tympanum large, distinct; dorsolateral 
fold sometimes present in young (large frog). 
R. magna Stejneger. 
cb Toes about two-thirds webbed; tympanum large; dorsolateral fold 
usually present (rather small frog) R. leytensis Boettger. 
6b Fingers and toes dilated into regular disks with a transverse groove 
around disk; no bony m.andibular teeth, 
cb A broad glandular dorsolateral fold; tympanum equals four-fifths 
eye; canthus rostralis rounded; toes about four-fifths webbed. 
R. erythrsea ISchlegel). 
cb A narrow dorsolateral fold. 
(f. First finger shorter than second. 
eb Snout truncate; finger disks large; tympanum three-fourths 
eye; canthus rostralis angular R. mearnsi Stejneger. 
eb Snout obtuse; t 3 Tnpanum about two-fifths eye; canthus rostralis 
angular R. dubita sp. nov. 
Smithson. Misc. Coll. 52 (1908-10) 437. 
