16, S 
Taylor: Philippine Amphibia 
261 
Variation. — Two small specimens taken in the same locality 
agree very well with the one described. One of them is red- 
dish brown above, the other grayish brown; the latter lacks 
the two spots on the breast, and the lip above is rather whitish. 
Two specimens taken at Concepcion, Busuanga, agree with the 
latter specimen in having a stripe on the upper part of the lores 
and the lip rather lighter. In these specimens the fold from 
the eye to the arm is very dim or wanting. 
Remarks . — Boettger described his Rana sanguinea in 1893, 
and Boulenger his R. varians in 1894. Both species are recorded 
as being related to R. temporalis Gunther. A comparison of 
descriptions reveals no essential differences. I am convinced 
that the two species are identical, and that the type of R. san- 
guinea Boettger is an immature specimen of R. varians Boulen- 
ger.' I have no specimens of R. sanguinea from the type locality, 
which is Culion, but have a specimen from a nearby locality, 
Busuanga, at a point on the island nearest Culion. Boettger 
describes a R. moluccana from Halrnaheira and Ternate, while 
in a later work he regarded this species as identical v/ith 
R, varians Boulenger. Barbour fails to agree with the later 
conclusion, but retains the species as distinct from R. varians 
Boulenger. Boettger evidently recognizes Boulenger’s species 
as distinct and makes no mention of R. moluccana as being 
related to R. sanguinea. 
Let us note the differences between the recorded description 
of the type and the description of the other two species. In 
Rana varians the tympanum is as large or nearly as large as 
the eye; in R. sanguinea it is more than three-fourths of the 
eye. In R. varians the interorbital distance is as broad as the 
upper eyelid or a little narrower; in R. sanguinea it is broader 
than a single eyelid. Rana varians has the toes nearly entirely 
webbed, the last two phalanges of the fourth toe free’; R. san- 
guinea has the toes three-fourths webbed, the last two phalanges 
of the fourth toe free. In R. varians the tibiotarsal articulation 
reaches the snout or beyond; in R. sanguinea it is one and one- 
half times as long. In R. varians the tibia is as long as the 
foreleg or a little shorter; in R. sanguinea it is slightly longer. 
None of the characteristics are contradictory; the only signi- 
ficant ones are the difference in the length of the hind leg and 
”Zool. Anz. 18 (1895) 132. 
”Abh. Senck. Nat. Ges. 25 (1900) 866. 
“Mem. Mus. Comp. Zool. Harvard Coll. (1912). 
