276 
Philippine Journal of Science 
1920 
and toes merely swollen or dilated in small discs; subarticular 
tubercles prominent ; a small oval inner metatarsal tubercle and a 
small round one at the base of the fourth toe ; tibiotarsal articu- 
lation reaches the eye or between the eye and the tip of the 
snout; back more or less distinctly granulate, with large flat 
granules at least on the sides.” 
Color . — “Olive or reddish brown, spotted and speckled with 
blackish ; lips dark with large whitish spots or bars ; limbs with 
dark crossbars ; lower parts whitish or buff, uniform or spotted 
with brown. Eye fieiy red. Male with vocal sacs on each side 
forming folds and a large oval gland on the inner surface of the 
arm. Snout to vent, 95 mm. Found in caves in total darkness.” 
Remarks . — The types were obtained in Sarawak by A. Everett, 
who later discovered the species in Palawan; I failed to find it 
in Palawan. Boulenger reports it from Malay Peninsula. 
Genus STAIJE.OIS Cope 
Ixalus GtJNTHEB, Cat. Batr. Sal. Brit. Mus. (1868) 15, part. 
Staurois Cope, Nat. Hist, Rev. (1865) 117. 
Rana BotTLENGER, Cat. Batr. Sal. Brit. Mus. ed. 2 (1882) 71, part. 
Micrixalus Boulengeb, Proc. Zool. Soc. London (1888) 205; Fauna 
Brit. India, Kept. (1890) 464. 
No vomerine teeth; tympanum distinct; fingers free, with 
disks; toes webbed; otherwise agrees very well with Rana.^^ 
One species is found in Palawan, Minadanao, and nearby 
islands. Very probably it does not enter Luzon, the western 
Visayan Islands, or Mindoro. 
“Boulenger, Ann. & Mag. Nat. Hist. IX 1 (1918) 374, has redefined 
this genus, listing five species: Staurois larutensis Boulenger, Borneo and 
Malay Peninsula; S. guttatus Gunther, Borneo; S. natator Gunther, Phil- 
ippines; S. nubilus Mocquard, Palawan; and S. tuberilinguis Boulenger, 
Borneo. He mentions the following generic characters: 
Tympanum small; with or without vomerine teeth; disks on fingers 
large, broader than long, larger than those of toes, with a half disk 
within the disk on the lower surface; toes fully webbed, the webs involving 
base of disks; outer metatarsal separated to base. 
The recognition of Mocquard’s species Ixalus nubilus = Staurois nubilus 
by Boulenger in this paper presents an opinion in which I do not concur. 
My own collection has more than one hundred specimens of S. natator 
from Mindanao, and I have recently been able to examine specimens of 
what is probably Mocquard’s Ixalus nubilis from various parts of Palawan 
and Busuanga. Occasional specimens from both lots show a papilla on 
the tongue; there appears to be no difference in average measurements; 
the granules on the Palawan specimens appear a little coarser than on 
most of the Mindanao specimens. 
