290 
Philippine Journal of Science 
1920 
Variation. — The large synonymy is evidence of a great amount 
of variation in this species. In Philippine specimens the color 
and markings vary greatly ; but relative measurements and pro- 
portions are rather constant, save in the case of the tympanum, 
which varies from one-half to nearly full size of the eye. 
The markings are usually in four to six longitudinal lines, as 
described above {Polypedates quadrilineatus) , or broken into 
numerous spots which do not conform to the outline of this 
design (P. maculatus) ; spots may be numerous or few, sometimes 
almost wanting. The ground color varies from very light yellow 
or whitish to dark purplish brown; sometimes (in life) almost 
black, with no markings visible. One of my specimens, taken 
in Mindanao, was bright lemon yellow over the entire body when 
first captured, with no markings. After being kept some time 
the yellow became grayish, and dim longitudinal stripes appeared. 
Wiegman had a Manila specimen collected by Meyer for the 
type of his Hyla quadrilineata. One of the types of Polypedates 
rugosus Dumeril and Bibron was from Manila- 
It is significant that the Polypedates maculatus form is rarely, 
if ever, found in Mindanao; in a collection of more than fifty 
specimens from Bunawan not one occurs. On the other hand, 
Manila specimens are largely of that type. 
Tadpoles. — Mandibles broadly edged with black, the upper 
forming a strongly arched, curved series, the lower a strongly 
V-shaped series; lower edge of lip with numerous papillae; upper 
labial extension with four series of mandibular teeth, the first 
uninterrupted, the second barely interrupted medially, the other 
two series very short, separated by mouth ; lower labial extension 
with three uninterrupted series. 
Color. — Yellowish brown to olive brown above, with black 
spot on nose and nostrils ; a bright yellow spot on point of snout, 
purplish on throat, lighter on belly ; tail olive, the crests colorless. 
It will be noted that these specimens differ from Flower’s 
description, in that in the form described above there are only 
four instead of five series of small teeth on upper lip, and that 
the three series on lower lip are uninterrupted. Flower also 
mentions a variation in a Singapore specimen. It may be pos- 
sible to differentiate the various races or subspecies of Polype- 
dates leucomystax by the characters of the tadpoles. 
Remarks. — The species occurs over the entire Archipelago; 
it has been reported from many localities in Luzon, and from 
Proc. Zool. Soc. London (1899) 899. 
