CEPHALOPODA OP THE HAWAIIAN ISLANDS. 
315 
Remarks. — The specimens which have been made the subject of the foregoing detailed description 
do not appear to offer any features in essential disagreement with Grant’s brief account of his Septa la 
stenodactyla from Mauritius, but with the recent more careful description by Hoyle “ of some south 
Pacific specimens which he referred to Grant’s species they are not in complete accord. The observed 
differences are admittedly trivial, but they maintain themselves persistently and with great constancy 
throughout the large series of specimens which has been examined and hence seem worthy of recogni- 
tion. As in the case of most closely related sepiolids the chief difference is in the structure of the hecto- 
cotylized arm. In Hoyle ’s figure of 5 . stenodactyla the modified papillae begin about halfway up the arm. 
Those of the ventral row are comparatively stout, fairly regular, not very closely appressed, and the 
figure shows only about fifteen of them. The conspicuous thickening of the arm at the point where the 
papillae begin and the strongly recoiled tip exhibited by the Hawaiian specimens do not appear. In 
E. scolopes, moreover, the modified papillae extend over a much greater proportional area (two-thirds) 
of the arm, are much more numerous (35 to 40 in the ventral row), and so tightly palisaded together 
that many of them are squeezed quite out of place. The details regarding the modified suckers of the 
remaining arms are also different from Hoyle’s description, especially in the case of the third pair, where 
in E. scolopes about twice as many of the suckers in the ventral series undergo enlargement. Unfortu- 
nately this comparison has been hampered by the lack of actual specimens of E. stenodactyla or any 
other Indo-Malayan representatives of the genus, so that the separate recognition of the Hawaiian race 
should perhaps be regarded as somewhat provisional. 
It is interesting to note that the differences separating the Japanese E. morsel from either steno- 
dactyla or scolopes are not very conspicuously greater than those just dwelt upon, but they appear equally 
constant, and I regard my original reference of the Hawaiian material to E. morsel as clearly erroneous. 
The males of the last-named species may be distinguished at a glance by the much more conspicuously 
enlarged suckers of the outer rows on the sessile arms, especially those of the second pair. Apart from 
secondary sexual characters, however, a description of one species, however detailed, would, so far as 
I am aware, serve almost equally well for either of the others, and I am at a loss to name any satisfactory 
criterion for the separation of any of these puzzling forms when represented by females alone. 
E. scolopes is one of the most abundant and ubiquitous cephalopods of the Hawaiian Islands. Not 
only is it a common surface form throughout the surrounding waters but it is to be captured on the 
reefs and even at considerable depths, as may be seen from the accompanying table. It is, however, 
a surprisingly constant species, the greatest variation observed being in the general form and propor- 
tions of the body, and even this is more probably due to the varying stresses of preservation than to any 
inherent differences in the animals themselves. It doubtless possesses luminous properties as glandular 
organs similar to those described by Meyer (1906) for Seplola, and more especially by Wtilker (1910, 
p. 26) for E. morsel are very^ conspicuous in a corresponding position within the mantle cavity. 
The specific name scolopes has been adopted in reference to the stockade-like appearance of the 
outer papillae on the hectocotylized arm. 
Subfamily STOLOTEUTHIN^E, new subfamily. 
Mantle and head united in the nuchal region by a commissure. Suckers in two rows, for the most 
part very small. Botlr dorsal arms hectocotylized, but the modification weak, consisting chiefly in 
the greater or smaller size and more crowded condition of the suckers. Eyes with a completely circular 
lidlike fold. Gladius, none. 
Genus STOLOTEUTHIS Verrill 1881 . 
Siolotculhis Verrill 1881, p. 417. 
Stololeuthis Verrill 1882, p. 375 [165] 
Body short and thick, bluntly rounded behind. Fins large. Arms short; all except tlie ventral 
pair united by a wide delicate basal web. Suckers for the most part minute. Mantle ornamented 
ventrally by a sharply delimited and specially pigmented shield-shaped area of the integument. 
Type.— -Seplola leucoptera Verrill 1878 (monotypic); a species of the New England region. 
Hoyle, 1904 , p. 24 , fig. B-D. 
