208 
MEMOIRS OF THE QUEEN SLANT) MUSEUM. 
The family Amphiprionidse, as here understood, includes three closely allied 
genera : Amphiprion (logotype, Lutjanus ephippium) , Actinicola (orthotype, Lutjanus 
percula), and Fhalerebus, gen. nov. (orthotype, Amphiprion okaUopisos Bleeker). The 
forms with conical or non-compressed teeth and less than 50 transverse series of 
scales on the body enter the family Chromidte. 
Premnas may be regarded as belonging to a separate family, the Premnidse, 
near the Amphiprionidse. Gheiloprion , Hem iglyph idodon , and Stegastes are also each 
probably worthy of family separation, to say nothing of several American genera. 
Although the order Amphiprioniiormes is easily separable from other orders 
of fishes, it is perhaps not very ancient, and most of the known genera appear to 
be emergent forms. Several fossils have been assigned to the “ Pomacentridse,” 
but none of them appears to enter the order as now understood. 
Priscacara Cope 2 from the Tertiary of the United States was at first compared 
with the Pomacentridse but Haseman 3 regarded it as <£ an ancestral Cichlid,” and 
Jordan 4 5 6 accords it family rank. I have, not seen the description of Chromis 
savorninip classed with the Pomacentridse in the Zoological Record. 
Odonteus Agassiz 0 was regarded as separable from the Sparoides and classed 
with the Scioenoides by its author, who remarked on its approach to Heliases. 
O. sparoides , the only species, is described as having large, well separated, conical 
teeth, first two dorsal spines subequal and less than half length of third, spinous 
dorsal high, body deep with a shallow caudal peduncle ; these features show that 
Odonteus is separable from the Amphiprioniformes. 
ARTIFICIAL KEY TO AUSTRALIAN AND SOUTH PACIFIC GENERA. 7 
A. Fifty or more transverse series of scales. D.ix-xi. Bases of spinous and soft dorsals of about 
equal length. All opercles serrated. 
B. Suborbital without a strong spine. AMPH1PRIONIDHC. 
C. Dorsals connected but notched. Occipital region naked. Caudal rounded 
Actinicola Fowler 1904. 
CC. Dorsals without notch. Occipital region scaly. Caudal truncate or excavate. 
D. Less than 18 dorsal rays. At least one tran verse band 
Amphiprion Bloch & Schneider 1801. 
2 Cope, Bull. U.S. Geol. Surv. Terr, iii, 1877, p. 816, and Yertebrata Tert. Format, i, 
1883, pp. 92 et seq. 
3 Haseman, Bull. Amer. Mus. Nat. Hist, xxxi, 1912, pp. 97-101. 
4 Jordan, Classif. Fishes (Stanford University Publications, Biol. Sci. iii, 2), 1923, p. 218. 
5 Arambourg, Nat. carte geol. Alger. (1) vi, 1927, p. 162, pi. xvi, fig. 4, and pi. xvii, fig. 3. 
6 Agassiz, Recherches poiss. toss, iv, 1839, p. 178. Not Odontceus Klug, Abhandl. Berl. 
Akad. 1843, p. 37 {fide Gemminger & Harold, Cat. Coleopt. iii, 1868, p. 1080), a genus of Coleoptera. 
7 Extralimital genera not dealt with here, but included in the order, are : Stegastes Jenyns 
1842, s.str. ; Fur car ia Pooy I860, preocc. = DemoiseUea Whitley 1928 ; Hypsipops Gill 1861 ; 
Microspathodon Gunther 1802; Euvchistodus Gill 1862; Pomataprion Gill 1863; Ayres! a Cooper 
1864; Onychognafhus Troschol 1866, preocc. Agripopa Whitley 1928 ; Hermosilla Jenkins & 
Evermann 1888 ; Nexilaris Jordan & Evermann 1896; Azarina Jordan & Evermann 1898; 
Ncxilosus Heller & Snodgrass 1903 ; Azurella Jordan 1919 ; Cenlrochromis Norman 1922 ; and 
Plectroglyphidodon Fowler & Ball 1925. Many of these may subsequently be removed to families 
or subfamilies distinct from the Pomacentrida?. Zahulon Whitley 1928 should be removed from the 
order on account of its three anal spines and superiorly situated lateral line. 
