320 
BULLETIN OF THE UNITED STATES FISH COMMISSION. 
orbital and supra-orbital lines, via tbe r. buccalis and r. ophthaloiicus superflcialis 
VII, respectively. 
IT nerve . — A somatic motor nerve. 
V nerve . — Tbe trigeminus contains viscero-motor and general cutaneous compo- 
nents. Tbe sensory fibers enter tbe spinal V tract, with wbicli is doubtless associated 
tbe chief sensory nucleus of tbe V nerve. Tbe Gasserian ganglion is clearly separable 
from tbe other ganglia of tbe Y + VII complex, tbougb most of tbe earlier writers 
on fish nerves have treated tbe whole collection together as the Gasserian. Cutaneous 
fibers go out from it to tbe r. ophthalmicus superflcialis V (which is fused with tbe 
nerve of tbe same name from tbe seventh nerve), tbe r. maxillaris V, tbe r. mandibu 
laris V, the truncus byomandibularis VII for the operculum, and, finally, the r. ophthal- 
micus profundus V. Tbe latter nerve consists of a few fibers which accompany the 
sympathetic radix longa of the ciliary ganglion to that ganglion, beyond which they 
can no longer be separately followed. Tbe relations of this nerve, which has not been 
reported by previous students of tbe teleosts, indicates that the embryonic profundus 
ganglion has fused witli the Gasserian. 
The motor fifth supplies tbe muscles usually described for teleosts, and in addition 
tbe so called m. genio-byoideus. Tbe innervation of this muscle has hitherto been 
usually assumed to come in teleosts from tbe VII nerve. This muscle is almost cer- 
tainly not homologous with tbe muscle iii the corresponding position of other verte- 
brates, which is innervated by tbe first spinal or bypoglossus nerve. 
Tbe IV and III nerves are purely somatic motor, vbile the II and I nerves can 
not as yet be placed in any of these categories. 
This investigation, which is still incomplete, has been carried out i)artly at 
Columbia University, jiartly at tbe Pathological Institute of tbe New York State 
Hospitals, and during tbe summer of 1898 at tbe laboratory of tbe United States 
Fish Commission at Woods Hole, Mass. Several morphological points which remain 
obscure can be finally cleared up oidy by tbe embryological method. Accordingly, 
during tbe present season I have secured and raised tbe eggs of Menidiu, putting up 
series of the embryos with a. view to some studies in tbe organogeny in tbe near future. 
Tbe eggs of our two common species (which I have elsewhere described) are 
very similar. Those of tbe larger species, Menidia notata, after artificial fertilization, 
can be easily reared in ordinary batching jars, and I obtained stages up to 6 days 
after batching (16 days after fertilization) and later stages by towing in tbe harbor at 
Woods Hole. But the eggs of the smaller species, Menidia gracilis, which inhabits 
tbe tide pools and protected bays, I was never able to batch, though tbe eggs seemed 
normal and fertile and tbe conditions were tbe same as for tbe other species. Tbe eggs 
Avould grow for from 6 to 12 hours and would then gradually die out. Possibly these 
fishes normally spawn in brackish water. 
I am under very special obligation to the United States Fish Commission, which 
has freely furnished all things needful for this work, and particularly to tbe director 
of its laboratory for many unusual courtesies. 
Granville, Ohio, Augtist 1, 1898. 
