308 
Fishery Bulletin 114(3) 
35 
30 
25 
20 
15 - 
10 - 
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25 - 
20 - 
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Brown rockfish 
Copper rockfish 
spnng 
fall 
uinler 
Figure 3 
Backtransformed mean counts (summed over 3 transects and all years) 
and 95% confidence intervals for brown and copper rockfish {Sebastes 
auriculatus and S. carinus, respectively) observed at Point Heyer Artifi- 
cial Reef , Puget Sound, WA, from summer 2005 through summer 2012. 
1 
] 
(95% ' 
; 
!' 97 % 
90% 
Brown rockfish >10 
Brown rockfish < 1 0 
Copper rockfish >10 
Copper rockfish < 1 0 
86% 
87% 
91% 
91% 
Spring 
Summer 
Fall 
Winter 
Figure 4 
Counts and percent proportions (summed over 3 transects and all years) 
of brown and copper rockfish {Sebastes auriculatus and S. carinus, re- 
spectively) <10 and >10 centimeters (total length) observed at Point 
Heyer Artificial Reef, Puget Sound, WA, from summer 2005 through 
summer 2012. 
[copper rockfish], P<0.05; two-tailed z-test for compari- 
son of 2 proportions; satisfactory n*Pi>5 and re[l-pi]>5 
sample size tests) (Fig. 6). 
Summed over all surveys, a total of 70 late-stage 
gravid brown and copper rockfish were observed, all 
of these in the spring and summer. Of the total num- 
ber of brown rockfish >10 cm observed in the spring 
(A(=377) and summer (N=998), 9% and 2%, respective- 
ly, were noted as late-stage gravid (we assume that no 
female rockfish reach maturity at <10 
cm [Washington et al.^; Gowan, 1983]). 
Late-stage gravid copper rockfish were 
only observed during the summer, and 
of the total number of copper rockfish 
>10 cm observed (493), 3% were noted 
as late-stage gravid (Fig. 7). If we as- 
sume a population sex ratio of 1:1, the 
percentages double with respect to the 
total number of potential female spawn- 
ers. In order to determine whether there 
were differences in time of spawning by 
length, we grouped the late-stage grav- 
id brown rockfish into 2 length classes 
(10<30 and >30 cm) on the basis of 50% 
maturity at approximately 30 cm TL 
(Love et al., 2002). The proportion of 
late-stage gravid brown rockfish >30 cm 
to the total numbers of late-stage gravid 
fish observed in the spring and summer 
was significantly greater in the spring 
(2=2.4, P<0.05; 2 -test for comparison of 
2 proportions; satisfactory n*pi>5 and 
re[l-pj]>5 sample-size tests). 
Fifty-one (7%) of the 718 brown 
rockfish >10 cm sampled by Hess et 
al. (2012) were fish that had been re- 
captured (2 of these were recaptured 
twice) according to genotype matching. 
The number of days at liberty between 
first and final capture ranged from 1 
to 1518 (M=615; SD=448.8). Thirty-one 
fish (4%) ranging in length from 17 to 
35 cm (M=27; SD=4.1), including the 2 
fish that were recaptured twice, were 
at liberty for more than one year be- 
tween first and final capture. A total of 
136 (19%) of the 718 samples were from 
late-stage gravid fish and 12 of those 
were captured twice in late-stage gravid 
condition in different years. The num- 
ber of days at liberty between captures 
for these fish ranged from 328 to 1469 
(M=635; SD=447.2). Overall, the lengths 
of the late-stage gravid brown rockfish 
sampled ranged from 21-35 cm (M=27; 
SD=3.28; includes only the length at 
time of first capture for fish that were 
captured twice). 
Miniumum and maximum recorded 
temperatures during 2006-2007 at the 
near- and offshore margin of the reef ranged from 7.5° 
to 14.6°C and from 8.1° to 13.4°C, respectively. Mean 
daily temperature changes were at least twice as great 
in the spring and summer as they were in the fall and 
winter at both locations, and daily temperature flucua- 
tions tended to be slightly greater at the nearshore 
margin year-round (Table 3, Fig. 8). The mean monthly 
air temperatures during the 12-month water tempera- 
ture recording period were all within 2% of the aver- 
