15 
Larval stages, for most species, unknown; adult in intestine of mammals, birds, and 
reptiles. 
Type genus. — Dibotliriocephalus Luehe, 1899. 
Of the six genera Dibothrioceplialus, Duthiersia, Scyphoceplialus , Botliridium, Diplo- 
gonoporus, and Pyramicocephalus, only two ( Dibotliriocephalus and Diplogonoporus , 
for key see p. 11) are reported for man. See also Sparganum, p. 83. 
Genus DIBOTHRIOCEPHALUS Luehe, 1899. 
Generic diagnosis. — Dibothrioceph alinee (p. 14): Scolex more or less elongate, 
groove-like suckers not highly developed. Beck present or absent. Genital organs 
in single series. Testicles and vitellogene glands in the lateral fields, occasionally 
anterior and posterior in the median field, in some cases reaching the median line. 
Vitellogene follicles, at least in part, situated between inner and outer longitudinal 
muscle layer. Uterus a long canal, in middle field, forming a rosette. Parasitic in 
man, felines, canines, pennipedia, and water birds. Larva (plerocercoid) in fish. 
Type species. — Dibotliriocephalus lotus (Linnaeus, 1758) Luehe, 1899. 
The 2 species of this genus which occur in man may be distinguished 
by the following key: 
KEY TO THE SPECIES OF DIBOTHRIOCEPHALUS REPORTED FOR MAN . a 
Head oblong; strobila 2 to 10 meters or more long, 20 mm. in maximum breadth; 
with 3,000 to 4,000 segments; the broad tapeworm D. latus (p. 15) 
Head cordiform ; strobila SO to 115 cm. long , not over 8 mm. broad; 400 to 660 segments 
present; found in Greenland; rare in man D. cordatus (p. 19) 
The Broad Tapeworm— DIBOTHRIOCEPHALUS LATUS & (Linnaeus, 1758) 
Luehe, 1899. 
[Figs. 1 to 13.] 
« Ward’s new bothriocephalid tapeworm, 1906. — Since this bulletin was sent to 
press, Ward has announced that he has obtained specimens of an apparently unde- 
scribed bothriocephalid tapeworm, taken from a six-year-old child, in the prairie 
region of Xorth America; the patient had eaten raw fish. Details will be published 
later by W ard. 
b Synonyms. — Teenia lata Linnaeus, 1758a; T. vulgaris Linnaeus, 1758a; T. grysea 
Pallas, 1761; T. membranacea Pallas, 1781; T. tenella Pallas, 1781; T. lata hominis 
Bloch, 1782a; T. vulgaris vel grisea vel membranacea Fischer, 1789a; T. lata vel Can- 
dida Fischer, 1789a; T. tenella Fischer, 1789a; T. vulgaris Gmelin, 1790a; Tenia inerme 
umana Brera, 1802a; Teenia inermis Weber’s Brera, 1803a [not Moquin-Tandon, I860]; 
Holy sis lata (Linnaeus, 1758) Zeder, 1803a; II. membranacea (Pallas, 1781) Zeder, 
1803a; Teenia humana inermis Budolphi, 1810a; Botryocephalus liominis Lamarck, 1816; 
Bothriocephalus latus (Linnaeus, 1758) Bremser, 1819a; Dibothrium latum (Linnaeus, 
1758) Diesing, 1850a; Bothriocephalus cristatus Davaine, 1873a; Teenia fenestrata Colin, 
1876a; Botrioceplialus vulgaris cristatus (Davaine, 1873) Grassi, 1880a; B. vulgaris latus 
(Linnaeus, 1758) Grassi, 1880a; B. vulgaris tenellus (Pallas, 1781) Grassi, 1880a; B. 
tenellus (Pallas, 1781) Grassi, 1880a; Bothriocephalus lalus tenellus (Pallas, 1781) 
Guzzardi Asmundo, 1885a; B. latissimus Burgnion, 1886; B. dorpatensis Thoma in 
Kuechenmeister, 1886c; B. balticus Thoma in Kuechenmeister, 1886c; Bothriocephalus 
latus fenestr[atus] Maggiora, 1891; B. fenestratus Maggiora, 1891; Dibotliriocephalus 
latus (Linnaeus, 1758) Luehe, 1899; D. cristatus (Davaine, 1873) Luehe, 1899. 
Bibliography. — Fo^ zoological discussion, see Leuckart, 1886, 864-929, figs. 357- 
393; for medical literature, see Stiles & Hassall, Index-Catalogue of Medical and 
Veterinary Zoology. 
