390 
M. G. Sykes 
interpreted as the split for the homotype; — au error iuto which 
my first examinatiou led me, but which was naturally dispelled by 
such cases as Figs 52 & 53. Pl. IX. It has been said that Miyake 
looked upou this longitudinal fission as a new appearance at this 
stage, but after the consideration of a complete series of preparations 
it seemed certain to me that this fission is the same as that seen 
in the spireme. I was therefore at first compelled to suppose that 
the Separation of the chromosomes along the previous line of fusion 
took place in the homotype and not in the heterotype division, while 
the transverse division which appeared to occur in the heterotype 
was another and quite different reduction division. This is some- 
what the interpretation given by Calkins 1 ), Stevens 2 ), Atkinson 3 ), 
Mottier 4 ), and it is of course impossible to bring it into line with 
other results which harmonise more perfectly with our present theories 
of reduction division. It may be possible that in some of these cases, 
as in Funlcia , division of the chromosomes in the heterotype does 
not consist in the Separation of two widely diverging longitudinally 
split rods, but in the division of the whole loop along a fission fol- 
lowing its whole length. 
X. Telophases and Reconstruction of daughter-nuclei. 
Fig. 56. Pl. IX. shows a telophase of the heterotype division, in 
which the longitudinal split is clearly visible in the chromosomes. 
This split can still be followed during the reconstruction of the 
daughter nuclei, (Fig. 57. Pl. IX.) and a paired structure is fouud 
even in the stage most nearly approaching a resting stage. (Fig. 59. 
Pl. IX.) A complete resting stage condition is never reached and 
no continuous thread appears-to be formed 5 ) but the nucleus here 
contains a number of chromatin masses, mostly double, united by 
threads of liuin. Fig. 60. Pl. IX. represents one of the prophases 
of the next division in which the split is well seen. 
c) The homotype division. The homotype division takes 
place at right angles to the last, and each chromosome, (of which 
six large and eighteen little pairs are present), is here generally 
attached to the spindle by one end, and divides longitudinally along 
the split seen in the anaphases and telophases of the last division, 
q Calkins. 1897. 
2 ) Stevens. 1898. 
3 ) Atkinson. 1899. 
4 ) Mottier. 1907. 
5) Miyake 1908. p. 112. also Gregoire and Wygaerts. 1903. 
