392 
M. G. Sykes 
vacuolated and is often seen to be connected witk one or more pairs 
of chromosomes 1 , 2 ). It is still present when tbe chromosomes are 
fully formed and does not disappear tili just about tbe time of tbe 
origin of tbe multipolar spindle. 
In tbe reconstruction of tbe daugbter nuclei, a single nucleolus 
was often seen, and Fig. 60. PI. IX. represents one of a few cases 
in whicb two or tbree nucleoli were present. In tbe young pollen 
grains tbere are also found two or tbree nucleoli, (Figs. 63. 64. PI. IX), 
but never more tban tbree. 
The facts detailed here appear to support tbe view 2 ) that tbe 
nucleoli are of value as a food störe for the nuclear reticulum and 
chromosomes. It is probably also of some interest that only two or 
tbree nucleoli are present after tbe reduction division, tbougb tbis 
miglit naturally be expected, since only half tbe amount of tbe so- 
matic cbromatin bas now to be stored in tbe nucleus. 
III. Somatic divisions. 
The somatic nuclear divisions have recently been described bv 
Strasburger, so that it would be superüuous for me to give a full 
description of tbe process and I propose to only shortly describe 
one observation wkicli appears to me to be of interest. 3 ) 
The striking nature of tbe double reticula seen in tbe very 
early prophases of reduction division led me to searcb for comparable 
pbenomena in the prophases and reconstruction of tbe nuclei in tbe 
preceding divisions. Similar appearances were commonly found in 
all tbe stages of tbe archesporial divisions; fully formed chromosomes 
were often seen lying together in pairs, and tbe paired nature of 
the knots, or so-called procbromosomes, whicb in Funkia 4 ) greatly 
exceed tbe somatic chromosomes in number, was obvious in most 
cases. These facts were particularly striking in the last archesporial 
division, tbougb it is impossible to say whether this is due to tbe 
greater size of the nuclei under consideration, or whether it bas any 
conuection witb a closer association of homologous cbromatin prepa- 
ratory to tbe fusion whicb will take place in tbe heterotypic spireme. 
In tbe ordinary somatic nuclei, seen in the cells of tbe petals, 
walls of the ovary, walls of tbe anther etc,, a paired structure is 
1) Cf. Wäger. 1904. 
2) Cf. Miyake. 1906. p. 101. 
3 ) Since the above was written, I have made a further examination of the 
somatic nuclei, the results of which will shortly appear in a note. 
4 ) See also Strasburger. III. 1906. pp. 7 & 40. 
