200 
Katharine Foot and E. C. Strobell 
zone B. In some forms he finds them between the large nuclei of 
zone B and in other forms on the periphery only. In Protenor it can be 
demonstrated that the small nuclei of zones A and C are astonishinglv 
alike and to this extent Protenor Supports Korsc.helt’s observations, 
but we find no evidence that the small nuclei of these two zones are 
continuous throughout zone B, therefore in this essential point Pro- 
tenor fails to support Korschelt’s interpretations. On the other hand 
the evidence Protenor offers in support of Will’s interpretation is more 
conclusive. 
It can be demonstrated that the large nuclei of zone B may fragment 
into a number of smaller pieces of varying sizes, and that these pieces 
may continue their development as independent nuclei, or again these large 
nuclear masses may appear sometimes as a single nucleus (photos 5 — 14), 
or may form a syncytium of small nuclei which are later set free — all 
these facts supporting Will’s observations on this point. The fragmenta- 
tion of a large nucleus into smaller pieces is shown in photo 25. The 
evidence indicates that nuclear fragments may develop as single nuclei 
without regard to their relative size, the larger pieces developing without 
further fragmentation, this fact perhaps offering one explanation why 
we find so many nuclei showing the same stage of development and 
yet differing so markedly in size. 
In the upper left corner of photo 25 is a large nuclear fragment which 
we believe is similar to the one shown in the right of photo 30 and in 
photo 31. Stages in the further development of such large nuclear frag- 
ments are shown in photos 26 and 27. Photo 32 demonstrates two smaller 
nuclear fragments showing the same stage of development as the larger 
pieces of photos 25, 30, and 31. Later stages of development are shown 
in photos 27 and 29. We believe these photos show in detail the phe- 
nomenon that Will found to occur at the base of zone B in the forms 
he studied. Although in Protenor it can be demonstrated that small 
nuclei may originate by fragmentation of large nuclei, we woidd by no 
means claim that all the nuclei of zone C have a like origin, for there is 
too mueh evidence of mitosis in this zone to warrant any such assumption, 
though the cases of mitosis are relativelv rare. 
The rare cases of mitosis as comparcd to amitosis have been demon- 
strated by Preusse (1895) in the ovaries of the forms studied by him. 
He questions amitoses being an evidence of degeneration of the cell and 
he believes in its functional significance. Gross (1905) also finds botli 
methods of division side by sidc in the ovaries of the insects he studied. 
That the features eharacteristic of amitosis are more or less obvious 
