The individuality of the chromosomes and their serial arrangement, etc. 127 
of growth such as is seen in the prophases of the somatic divisions. The 
long axes of the homoeotypic spindle figures may lie at any angle with 
reference to each other and they may lie in almost any position with 
reference to the apex of the pollen mother-cell. Figure 30 shows a 
cell in which the two spindles are at right angles to each other with one 
lying nearer the apex of the mother cell. This figure shows both a lateral 
and a polar view of the equatorial plate. The former shows an edge \äew 
of a rather flat plate. The polar view makes it possible to count the 
thirty-seven individual oval chromosomes and there is evidence that 
here again they are connected together in a series. In such ^üews as 
are shown in figure 30 the series can be foUowed throughout most of 
its length. The behavior of the chromosomes in these homoeotypic 
divisions is quite similar to that which I have described for the somatic 
divisions. The spindles are about one-half the size of the heterotypic 
spindle figure. In the telophases the remains of the central spindles 
are still present, but not strongly developed, nor do they persist and 
form cell plates. No central spindle figure or cell plates between the 
granddaughter nuclei appear in my preparations. 
The Development of the Pollen. 
Three of the four daughter nuclei, as described by Juel for Carex 
acuta, now migrate toward the apex of the spore mother-cell and become 
crowded togethei^. As shown in figure 31 these nuclei are small. They 
contain for a time conspicuous nucleoles and the chromosomes are clearly 
defined oval masses distributed about the periphery. The fourth nucleus 
lies near the center of the general cytoplasmic mass. It is larger than 
the others, especially as it prepares for the division which forms the 
vegetative and the generative nuclei. The masses of chromatin which 
are present at this stage are difficult to count with absolute exaetness, 
but the number is never larger than that of the haploid chromosomes. 
Juel (1900) describes for Carex acuta the development of the gene- 
rative cell by a sort of free cell formation which, however, judging from 
his figure is quite different from the method of free cell formation in 
the embryo sac of Ephedra altissima as described by Strasburger (1880). 
In Juel’s figure the oell boundary is on the inside of an apparently fibril- 
lär Zone. The method of formation of the generative cell, as described 
by Juel, is furthermore at variance with what described by Strasburger 
(1884) for Heleocharis, and with what is considered to be the common 
method in Angiosperms. Friemann (1910) has recently studied the 
development of the generative cell in a number of monocots {Fritilhria 
