A critical study of the cytolog\' of Crepis virens. 
99 
Allen (1) described chromatic aggregations in the presynaptic 
pliases of Lilium canadense, but ascertained that they are more numerous 
than the chromosomes. A similar conclusion was reached by Miyake (33). 
Gregoire (20) exaniined Lilium martagon and his results agree with 
those of Allen; Gregoire does not consider the chromatic aggrega- 
tions to be true ‘ganiosomes’. 
Miyake (33) observed that m the heterotype prophases of Galtonia 
candicans, Allium Victorialis, Tradescantia virginica and various species 
of Iris that the nnmber of the paired chromatic bodies, or ‘zygosomes’ 
approximated to that of the rediiced nnmber of chromosomes. On the 
other hand in Lilium the nnmber of bodies appeared to be greatcr, whilst 
in Funkia sieboldiana it appeared to be less than that of the specific num- 
ber of chromosomes. Sykes (56) also exaniined the early heterotype 
prophases of Funkia sieboldiana, and described ‘knots’ composed of dark 
granules embedded in a colourless substratum. She was unable to count 
the knots accurately, but found that their nnmber far exceeded that 
of the somatic chromosomes. 
Gregoire (20) studied the heterotype prophases of Allium fistu- 
losum, and his results do not corroborate those of Miyake. Gregoire 
recognised two types of presynaptic phases, in the one type chromatic 
aggregations were far more developed than in the other. He does not 
consider the ‘noeuds’ to be ‘gamosomes’ as the chromatin is not entirely 
relegated to them, but is partially distributed as granules in the reticulum. 
Tischler (60) described the chromatic contents of the resting pre- 
meiotic nuclei in Bryonia as “eine Ansammlnng von Körnchen an be- 
stimmten Punkten, die Ähnlichkeit mit den OvERXON’schen Prochromo- 
somen aufweisen. Sie sind aber, so weit ich sah, wechselnd an Zahl und 
Größe, und ich vermag nicht anzugeben, ob schließlich jedes Mal genau 
alles Chromatm in diesen Punkten aufgeht” (p. 85). 
Mottier (35) could not determine the exact nnmber of the chromatic 
masses in Podophyllum peltatum and moreover found that they varied 
both in size and in degree. 
Eosenberg has extended the investigation of resting nuclei over 
a wide ränge of plants, and has contributed his results in a series of in- 
teresting papers (44, 45, 46, 47 and 48). He has shown that the restmg 
somatic nuclei of niany plants possess chromatic bodies which correspond 
approximately in nnmber to that of the chromosomes. In the pre- 
synaptic prophases these bodies pair, in preparation for the formation 
of the bivalent chromosome of the heterotype division. He has adopted 
the name ‘prochromosome’ for these bodies, though he is of opinion 
1 * 
