100 
L. Digby 
that tliey woiüd have beeil more correctl)" designated as cliromosome 
‘centres’ (47). 
OsTENFELD aiid lie have also shown that in Hierammi auricula 
and in H. venosum (39) the number of the prochroinosomes approxiniates 
to that of the chroniosomes. In Crepis virens (46) Rosenberg was able 
to check the nuinerical identity of chromosomes and prochromosonies, 
and States that “die Anzahl der Prochroniosonien beträgt in wirklich 
ruhenden Kernen 6, in den Pollenzellkernen 3” (p. 75). 
This is not in accordance with Beer’s observations. In a receiit 
paper (3) on spore development in the Compositae, in which Crepis virens 
is included, he States that, “durmg the period just preceding sjmapsis 
the nuclei of those Compositae which have been examined were found 
to contam a more or less fine reticuluni. No definite aggregations of 
chroniatin which could be regardcd as prochromosonies Avere in any 
case found upon this reticuluni” (p. 722). 
According to Rosenberg (47) the number of the prochroinosomes 
in the vegetative restmg nuclei of Xuphar luteum and of Pinguicula vul- 
garis is frequently less than that of the chromosomes, and he attributes 
this fact to an amalgamation of several prochroinosomes. In the pre- 
synaptic stages of Drosera longifolia, D. rotundifolia and the hybrid 
D. ohovata, the prochromosonies and the chromosomes coincide in theK 
respective numbers. 
Lagerberg (25) described honiogeneous deeply staining bodies in 
the restmg enibryo sac mother-cell nucleus of Ädoxa moschatelUna cor- 
responding to the gamosomes of Strasburger, biit he was not able to 
correlate their number AA'ith that of the chromosomes. 
Yamanouchi (61) has ascertained that ‘prochromosonies’ are present 
in restmg nuclei throughout the life history of Pohjsiphonia violacea, in 
about the same numbers as the chromosomes. He describes the resting 
nucleus of the germmating tetraspore as possessing a delicate reticuluni 
dotted AA'ith chroniatic granules. Prior to niitosis the iietAvork beconies 
coarser, and the granules collect to form chains of beads of Awyüig 
lengths. These are ‘prochromosonies’. They gradually Iiecome more pro- 
nounced, and theh- bead-like structure is transformed into the more 
honiogeneous rod shaped chromosomes. 
ÄIooRE and Embleton (34) haA'’e found that chroniatic rods are 
present in the resting nuclei of Triton and that their number generally 
corresponds to that of the chromosomes. These rods pair to form the 
gemini in the heterotype prophases. “The chief interest of these bodies 
lies in the fact that they obAÜously represent the chroniosomes of division 
