106 
L. Digby 
sliarper outline. The chromatic bodies, witliin the nucleus, coustantly 
exhibit a paired arrangement (figs. 4, 5, 6, 7, 8, and 9), and the sanie 
phenomenon may also be observed in the extra nuclear bodies scattered 
in the cytoplasm (figs. 4, 5, 6, and 7). It is beüeved that tliis pah’ing 
represents an early fission for the next .mitosis. The origm of the 
pahed arrangement is difficult to determine owing to the fact that the 
chromatic bodies are evolved by gradual chromatic concentration. It 
is probable that the pairing may somethnes be due to an early association 
of separately Condensed daughter segments, but as a rule it appears to 
be the result of fission of a segment Avliich has Condensed as a Avhole. 
In the upper nucleus of fig. 4, the chromatic granules are evidently as- 
sociatmg, whilst in the lower nucleus, and m fig. 5, fission seems to be 
takmg place. 
The chromatic bodies have been counted m the, relatively speaking, 
‘restmg’ nuclei that have been found in the archesporial tissue. The 
number of these bodies is small, probably it does not exceed six, the 
somatic number of the chromosomes, but the number is variable and 
inconstant. It is not advisable to attach too inucli importance to the 
variability in numbers of the chromatic bodies within the archesporial 
nuclei, for, as these nuclei are not in complete ‘rest’, they do not lend 
themselves to critical examination. Kosexberg (45) has expressly stated 
that ‘prochromosomes' are most constant in number in the resting nuclei 
of mature tissues (45, p. 402). This condition of ‘rest’ can hardly be 
applied to the young and quickly dividing archesporial cells. Resting 
nuclei of the tapetum have therefore also been studied, and the re- 
sults of those observations ^Yill be given at the end of the description 
of the archesporial niitoses. Moreover, though the typical bodies stand 
out distmctly from the other nuclear contents, yet smaller chromatic 
aggregations and particles present in the nucleus, afford controversial 
points as to their nature. Should they be regarded as chromatic bodies 
or as portions of chromatic bodies, and hence be included in the counts ? 
Or are they merely nucleolar fragments, or independent chromatic 
granules? Davis (6 and 7) has emphasized this difficulty in considering 
the counts of chromatic aggregations in restmg nuclei of sonie Oenotheras. 
Figs. 5—9, inclusive, are drawings taken from uncut nuclei m the 
coneütion of so-called ‘rest’ which intervenes between successive archespo- 
rial mitoses. 
The study of these nuclei strongly suggests that the chromatic bodies 
present in the nucleus are concentrations, or storehouses of chromatin 
(Laibach 26, Rosexberg 47, Lundegardh 29), whose substance will 
