L. Digby 
1 22 
Metaphase. 
A polar view of a homotype equatorial plate (fig. 92) sliows the three 
concentratecl chromosomes to be more or less V shaped, due to the partial 
cleavage of the daughter chromosomes. As they pass to the poles the large 
chromosomes are constantly hooked (fig. 93) (cf. Rosexberg 46, Taf. 1, 
figs. 25 and 26). It is possiblc that this appearance may not always be 
due to a bending over of the chromosome, but that it may represent 
the opening out of the fission which is visible in the subsequent retro- 
gressive telophase stages (fig. 95). 
Anaphase. 
Arrived at the poles, the chromosomes mass together in the usual 
characteristic way (fig. 94). x\s they separate out once more, they tend 
to break apart transversely. Consequently, in late anaphase, there fre- 
quently appears to be a larger number of chromosomes present than 
the three typical of the reduction divisions. Rosexberg (46) has 
emphasized this fact and has figured these stages. He suggests that 
this evident loose adhesion of the units which go to build up a chronio- 
some, may indicate that the chromosomes of C. virens are of a collective 
character, that is to say that they may represent associations of chromo- 
somes. This Suggestion is further corroborated by the observation that 
the portions of the several chromosomes remain unconnected until, as 
concentrated clmomosomes, they take up their position on the equatorial 
plate. It is only therefore at metaphase, and as they proceed to the poles, 
that they are distmet entities. 
Telophase. 
Great variety is exliibited by the nuclei as they pass through the 
retrogressive phases into rest. Sonie of the chromosome portions may 
remain more or less entire as large rounded bcads (fig. 96), suggesting a 
previous transverse Segmentation of v'hole chromosomes, vliile other 
portions may show fission in their substance (fig. 95), with paired threads 
joining the sides of one split portion to those of another (fig. 95). 
Rosexberg (46, Taf. 1, fig. 29) has figured beautiful examples of 
fission in the chromatic contents of the telophases of tetrad nuclei, and 
he conceives it to be the fission preparatory for the next nuclear division 
(46, p. 72). 
As the tetrads are about to separate from one another (fig. 96) the 
chromatic nuclear contents are distributed in the form of rounded beads 
