126 
L. Digby 
(20, PI. 2, fig. 37) which passes iiito synapsis, the otlier, a inore clevelopecl 
rcticuluni with larger chroiuatic aggregations (20, PI. 2, fig. 36). The 
same type of synapsis ensues from botli series. x\lso in the presynaptic 
stages of Liliion martagon he has shown that the ‘plages’ of chroinatin 
are of different sizes and shapes, and in sonie nuclei they may be niore 
developed than in others. 
Mottier (.35), working on the same plant, describes a siniilar diver- 
gence m the character of the propliases, and States that this difference 
also embraces the synaptic phase. A finely granulär synapsis (35, PI. 28. 
fig. 21) results from the series of propliases whose chromatin is distri- 
bnted in more or less imiformly sized granules, whilst a relatively lumpy 
synapsis (35, PI. 28, fig. 22) results from the series of propliases whose 
chromatin is in large irregulär masses. He has also found two types of 
propliases in the resting vegetative and pollen mother-nuclei of Podo- 
phylhm peltatum. In the resting pollen mother-nuclei the chroinatin 
may be in medium sized granules (35, PI. 27, fig. 1) or in large lumps 
(35, PI. 27, fig. 2). In early presynaptic prophases, nuclei, even in the 
same locuhis, may exhibit diverse characters. In one, the chromatin 
may be aggregated into irregulär shreds (35, PI. 27, fig. 3) whilst in an- 
other, it may be distributed throughout the linin as small granules (35, 
PI. 27, fig. 4). Subsequently Mottier (36) has extended these obser- 
vations and has experimented in fixing the embryosac mother-cells of 
Lüium martagon and Lilium candidum. He cut away the waUs of the 
ovaries, in sonie cases leaving the cavity enclosed by a thin layer of 
tissue, whilst in others the cavity was laid bare. When the fixmg fluid 
had to penetrate a tissue, six or seven layers of cells in thickness before 
it reached the ovule, the chromatin of the embryosac mother-cell nu- 
cleus was found to be precipitated in large angular lumps (36, PI. 23, 
fig. 2) whereas when the fluid had direct access to the ovule, the chromatin 
of the embryosac mother-cell nucleus was precipitated in fine granules (36, 
PI. 23, fig. 1). Mottier ascribes this difference to the action of the fixing 
fluid. If the fluid be rendered weaker owing to its passage through several 
layers of cells it caiises a coarser precipitation of chroinatin, whereas if 
its strength be unimpaired a relatively fine precipitation ensues. 
]\Ieves (32) has figured resting spermatogonial nuclei, fixed in Her - 
5IAXXS fluid, from different portions of the testis of Salamandra maculosa. 
The nuclei of the anterior lobe show their chromatin to be somewhat 
massed into irregulär aggregations, ‘Chroniatinbrocken ’(32, Taf. 1, fig. 4) 
whilst the nuclei of the extreme tip show a fine chromatic precipitation 
(32, Taf. 1, fig. 5). 
