272 
Sidney 1. Kornhauser 
the chromosomes ^Yhicll in early prophase were sometimes connected to 
form the large ring-tetrad (Plate XX, Figs. 110 and 111), but later, 
as the tetrad shortened, puUed apart (Plate XX, Figs. 113, 115). The 
first spermatocyte division separates "whole chromosomes, which are 
joined end to end to form gemini in both E. hinotata and E. curvata. The 
division, being transverse, is reductional and corresponds to the hetero- 
typic division of such forms as often retain the ring form of tetrad in 
the first division after parasyndesis, as described by the Schreiners 
(’06a) for Toinopteris and by Janssens et Willems (’09) for Älytes. 
In ÄmpMscepa (p. 267, Figs. B— H) both ends of the large ring-tetrad 
often remain connected; the insertion of the spindle fibers is lateral, not 
terminal, and the first division, though reductional, separates two chromo- 
somes which lie parallel to each other with their long axes transverse to 
that of the spindle. 
In E. hinotata, the x- and ?/-components of the allosonie pair separate 
froni each other reductionally and neither lag behind nor precede the 
autosomes in division. In E. curvata, the a;-element normaUy passes to 
one of the two daughter cells undivided. It is always lengthened in the 
direction of the spindle owing to its attachments to the separating auto- 
somes and is sometimes divided into two parts, although this phenomenon 
is unusual and probably abnormal. 
IX. Division of the Second Spermatocyte. 
A. Enchenopa binotata. 
There are ten chromosomes in all the second spermatocytes (Plate 
XXI, Figs. 164—168). The macrochromosonie is broad (Fig. 169) and 
not elongated in the axis of the spindle. IMien seen in end view (Figs. 164 
to 166) it therefore appears larger, in proportion to the other chromo- 
somes, than it did in the first maturation division. The shape of the 
macrochromosonie in this division is corrclated with its longitudinal 
division, which was already seen in Figure 141 (Plate XX). 
There is nothing unusual about the second division. The chromo- 
somes separate concomitantly (Plate XXI, Figs. 170—172, 174), and 
are weil separated from one another lateraUy, so that it is possible to 
count late anaphases with perfect accuracy (Plate XXI, Fig. 173). In 
such anaphase plates and in spermatids from smear slides (Plate XXI, 
Fig. 175, 176, 177) there were always ten well defined chromosomes. 
I found no evidence of an unequal distribution of the chromosomes 
