284 
Sidney I. Kornhauser 
The Symbols, black for “autochromatin”, white (i. e. ontlines only) for 
“ic-chromatin”, small cii’cles for “ 2 /-chromatin” and dots for liniii, are of 
coui’se only hypothetical. Figure 200 (Plate XXII) represents the 
bouquet stage of E. Unotata. The autochromosomes are paired side by 
side, the allosonies {x and y) have united end to end. The a;-component 
contains “a>chroniatin” and “;y-chromatin”, the ^/-component contains 
only “y-chromatin”, but equal in amount to that of the a:-component. 
Figure 201 shows the lateral approximation of the aUosomes, diie 
possibly to the attraction of “^-chromatin” for y-chromatin. The auto- 
somes have given up their polar arrangenient. In the strepsistene stage 
(Fig. 202) the aUosomes separate, the a;-component shortens and the y- 
component appears less granulär. The paired autosomes separate along 
their lengths and twist about each other. In Figure 203 the aUosomes 
have contracted still farther, the a;-component more so than the y- 
component, and they show a thick connecting Strand between them. 
The autosomes have contracted and split longitudinally. Figure 204 
represents a stage slightly older than Figure 203 and shows a complete 
condensation of the aUosomes and the straightening out of the tetrads. 
FoUowing this is the metaphase of the first spermatocytedivision (Fig, 205). 
This is a reduction division for both the autosomes and the aUosomes. 
The rc-component passes to one of the resulting cells (a, Fig. 206), the 
?/-component to the other ceU (b, Fig. 206). The divisions of the second 
spermatocytes are represented in Figures 207 and 208. The division of 
cell a gives rise to two spermatids (Fig. 208 a^ and a^) of like compo- 
sition. Each contains half the substance of the o^component of the 
allosome pair and is “female determining”, having both “ä- chromatin” 
and “y-chromatin”. CeU b gives rise to two “male determining” sper- 
matids (Fig. 210 bl and b^), which contain no “a>-chromatin”, but 
chromatin” equal to that of a^ and a^ (Fig. 208). 
Turning now to E. curvata (Figs, 211—222), the bouquet stage is re- 
presented in Figure 211. It shows two paUs of autosomes (one the macro- 
chromosonie pair), the a;-element (composed of “x-chromatin” and a 
smaU amount of “^/-chromatin”) situated at the positive pole of the 
nucleus, and the chromatic nucleolus, which is in intimate relation with 
the a:-element. It is assumed that this nucleolus, which stains like an 
aUosome, is composed of “^/-chromatin”, part of which originaUy be- 
longed to the rc-element, and that when the autosomes passed froni the 
fine network stage (foUowing the last spermatogonial division) into the 
early leptonema and began to pair, this “«/-chromatin”, stül attracted 
by the “?/-chromatin” of the a;-element, gathered at the positive pole. 
