158 
Katliarine Foot and E. C. Strobell 
Photo 11 shows a germinal vesicle of a relatively large egg neai' the 
distal end of the ovary (text-fig.). The eggs at the extreme distal end 
are much larger and many of the germinal vesicles of these eggs are fnUy 
as large as those of the eggs foimd in the receptaculum ovornm. 
After leaving the ovary the eggs are deposited in the receptaculum 
ovorum where they must remain from one to three days, as we found that 
these worms deposit a cocoon not oftener than every third day [Foot and 
Strobell (05)]. 
ülany of our Photos of the germinal vesicles of the ovarian eggs 
show them to be sometimes even larger than those of Photos 12 to 18 
which are from eggs found in the receptaculum ovorum. We believe there- 
fore, we are justified in stating that the eggs of Allololophora have reached 
tlieii- maxhnum size before leaving the ovary. AYe are further justified 
in stating that the evolution of the second group of reduced chromosomes 
does not begin until after the eggs have completed theii’ “grand 
accroissement” for the leptotene, paehytene and diplotene stages 
occur in eggs found in the receptaculum ovorum and it is only exceptionaUy 
that the largest eggs ot the extreme distal end of the ovary have developed 
even to the leptotene stage. On the other hand the majority of eggs in 
the receptaculum ovorum have not yet developed to this stage. 
Photo 12 shows the earliest stage of development of these eggs. The 
chromatin is evenly distributed throughout the entire germinal vesicle and 
there is no indication whatever of the chromatin threads 
characteristic of the leptotene stage. In this preparation both 
the larger and small nucleolus are present. 
Photo 13 shows one of the earliest phases of the leptotene stage. AA"e 
have many preparations showing the threads still more delicate — too 
delicate to admit of a satisfactory demonstration in a photograph^). 
In Photo 13 the large nucleolus is broken by the technique and covers 
a larger area than is usual. The small nucleolus is probably obscured by 
the larger, as they are often in dose contact as in Photo 15. This is seen 
also in Photos 118, 124 and 125 Foot and Strobell (05). 
ln Photo 14 the leptotene threads are somewhat thicker and some of 
the Strands show a longitudinal split, but we Interpret this as forecasting 
the longitudinal split so j)ronounced in the later stages. E. g. Photos 16, 
17 and 19 [sce also Foot and Strobell (05) Photos 114, 115, 117, 119, 120, 
121. 123, 124 and 126], 
1) Other examples of the leptotene stage in these eggs are demonstrated in 
Photos 111 and 113 Foot and Strobell (05). 
