232 
J. F. Mc Clendon 
reduction takes place, whether the egg be parthenogenetic or not. Shall 
this fact prove universally true Weismaxx’s hyiiothesis is erroneous and 
the question of pre- or post-reduction can have no significance in here- 
dity. For this reason we will not discuss this question in relation to the 
spermatogenesis of Pandarus. The significant fact is that reduction has 
taken place, i. e. that each spermatid contains numerically one half of 
the same individual chromosomes that were present in the primary 
spermatocyte. This process is shown diagrammatically in the accom- 
panying text figure. 
Oogenesis. 
The oogenesis is similar to the spermatogenesis tlirough the syne- 
zesis Stage. The primary oocytes are larger than the primary spermato- 
cytes when first formed, and this differcnce increases as growth proceeds. 
The leptotane stage is shown in fig. 14, and two cells in the synezesis 
Stage are seen in fig. 15. Immediately after synezesis a plasmosome is 
found, but whether it existed during synezesis coiüd not be determined 
owing to the massed condition of the clmomosomes. 
Eight filaments radiate from the plasmosome and in favorable as- 
pects are seen to be double, thus showing that the diplotane stage has beeil 
reached, fig. 16. These eight double filaments arose by a longitudinal 
synapsis in the synezesis stage, from the sixteen single filaments of the 
leptotane stage. 
The growth period now begins, fig. 16; the cell outlines first disap- 
pear but reappear some time after the oviduct is reached. During the 
syncytial stage the cytoplasm becomes filled ivith basophile gTanules. 
.MoROFpi) found, in the eggs of free swimming copepods, basophile gra- 
nules extruded from the nucleus into the cytoplasm, but I have not noted 
a similar origin of the granules mentioned above. The oocytes have now 
become much flattened and suggest in their arrangement a pile of coins. 
The yolk is deposited in the form of proteid bodies ond oil droplets. The 
plasmosome becomes vacuolated and is probably in process of solution. 
Fig. 17 shows a nucleus and fig. 18 a section of an entire oocyte on a 
less magnified scale, at this stage. The black bodies in the cytoplasm 
are proteid and the light sjiots are fat. 
The yolk accumulates and crowds the nucleus. The nuclear sap be- 
comes basophile and the nuclear wall disappears leaving the eight double 
filamentous chromosomes radiating fiom the large plasmosome and sur- 
1) Arch. f. Zellforschung, II, S. 432. 
