408 
GOMPHRENA 
Gomphrena Linn. Amarantaceae (4). 90 sp. S. Am., Austr. Herbs 
with cymose heads of firs. Fir. £ with 5 hairy perianth-leaves and 
5 sta. united into a tube. 
Gonatanthus Klotzsch. Araceae (vi). 2 sp. Himal. 
Gongora Ruiz et Pav. ( Acropera Lindl.). Orchidaceae (19). 20 sp.trop. 
Am. Epiphytes with hanging firs, whose ovary is so bent that the 
labellum comes to stand above the column. The sepals and petals 
spring from the column (an argument for its axial nature). See 
Darwin, Orchids , p. 166. 
Goniolimon Boiss. = Statice Tourn. 
Gonolobus Michx. Asclepiadaceae (11. 5). 70 sp. Am. 
Goodenia Sm. Goodeniaceae (1). 70 sp. Austr. 
Goodeniaceae. Dicotyledons (Sympet. Campanulatae). 12 gen. with 
210 sp , chiefly Austr., a few N.Z., Polynes., and trop. coasts. Herbs 
and shrubs with alt. exstip. leaves and no latex. Firs. $ , zygo- 
morphic, solitary in the leaf-axils or in cymes, racemes, or spikes. 
K usually 5, small; C (5); A 5, alt. with the petals, epipetalous or 
not, with introrse sometimes syngenesious anthers ; G (2), inferior or 
semi-inferior, rarely superior, 1- or 2-loc. ; style simple with ‘pollen- 
cup ’ close under the stigma. Into this the pollen is shed in the bud ; 
“the cup then closes up, leaving only a narrow opening for the most 
part covered by hairs. At the same time the style bends down to 
stand in the mouth of the almost horizontal flr., so that insect-visitors 
come in contact with the hairs and dust themselves with a little of the 
powdery pollen. As the stigmatic lobes grow up in the cup they 
keep forcing fresh pollen into the narrow slit, and finally emerge by it 
themselves and then receive the pollen of younger firs, from insect- 
visitors (Muller). The mechanism should be carefully compared 
with that of Campanulaceae and Compositae. Ovules usually ascend- 
ing, anatropous. Fruit usually capsular, sometimes a nut or drupe. 
Embryo straight, in fleshy endosperm. 
Classification and chief genera (after Schonland): The G. are very 
closely allied to Campanulaceae, differing chiefly in the absence of 
latex and the presence of the pollen-cup. They resemble Gentian- 
aceae in a few points. 
I. GOODENIOIDEAE (firs, rarely in heads; corolla folded 
in bud; ovary inf. or J-inf., with usually 2 or more ovules ; 
endosperm): Velleia, Goodenia, Leschenaultia, Selliera, 
Scaevola, Dampiera. 
II. BRUNONIOIDEAE (firs, in heads; corolla valvate; ovary 
sup., i-ovuled; no endosperm): Brunonia (only genus). 
[Placed in Campanales by Benth. -Hooker, in Campanulinae by 
Warming.] 
Goodenovieae (Benth.- Hooker) = Goodeniaceae. 
Goodia Salisb. Leguminosae (in. 3). 2 sp. S. Austr. 
Goodyera R. Br. Orchidaceae (4). 25 sp. N. temp., trop. As., New 
