RUBIACEAE 
5^5 
C (4 — 5), epigynous, valvate, convolute, or imbricate ; A 4 — 5, 
alt. with petals, epipetalous ; G (2) rarely (1 — 00), 2-loc. with 
1 — 00 anatropous ovules in each loc. ; ovules erect, pendulous, or 
horizontal; style simple ; stigma capitate or lobed. Fruit a capsule 
(septi- or loculi-cidal), berry or schizocarp. Embryo small, in rich 
endosperm. 
Most R. have conspicuous insect-pollinated firs. The Brit. sp. 
have small firs, with freely exposed or but slightly concealed honey, 
and are chiefly visited by flies (see p. 89) ; of the trop. sp. many have 
bee- and Lepidoptera-flrs. with long tubes. Honey is usually secreted 
by an epigynous nectary round the base of the style. Heterostylism 
is common, and dicecism sometimes occurs. [See Burck in Ann. 
Buitenz. III. 1883, and iv. 1884.] 
Several R. are of economic importance, eg. Cinchona, Coflea, 
Uragoga, Rubia, &c. 
Classification and chief genera (after Schumann). The R. are 
closely allied to Caprifoliaceae (q.v.) and less nearly to Compositae, &c. 
I. CINCHONOIDEAE (ovules 00 in each loculus). 
A. Cinchoninae (fruit dry) : 
a. Firs, solitary or in decussate panicles. 
a. Fir. regular ; seed not winged ; corolla valvate. 
1. Condamineeae : Condaminea. 
2. Oldenlandieae : Oldenlandia, Houstonia, Pentas. 
b. As a, but corolla imbricate or convolute. 
3. Rondeletieae : Rondeletia. 
c. As b, but seed winged. 
4. Cinchoneae: Cinchona, Bouvardia, Cosmibuena. 
d. As a, but corolla 2-lipped. 
5. Henriquezieae : Henriquezia. 
/ 3 . Firs, in heads. 
6. Naucleeae : Uncaria, Nauclea. 
B. Gardeninae (fruit fleshy) : 
7. Mussaendeae (corolla valvate) : Mussaenda. 
8 . Gardenieae (corolla imbricate or convolute) : Randia, Gar- 
denia, Posoqueria, Duroia. 
II. COFFEOIDEAE (ovules 1 in each loculus). 
A. Guettardinae (ovule pendulous ; micropyle facing upwards) : 
9. Alberteae: Alberta. 
10. Knoxieae: Knoxia. 
11. Vanguerieae : Plectronia, Cuviera. 
12. Gziettardeae : Guettarda. 
13. Chiococceae : Chiococca. 
B. Psychotriinae (ovule ascending; micropyle facing down- 
wards) : 
a. Corolla convolute. 
14. Ixoreae : Coflea, Ixora, Pavetta. 
