RUB I ACE AE 
549 
have small firs. with freely exposed or but slightly concealed honey, 
and are chiefly visited by flies (see p. 89) ; of the trop. sp. many have 
bee- and Lepidoptera-flrs. with long tubes. Honey is usually secreted 
by an epigynous nectary round the base of the style. Heterostylism is 
common, and dioecism sometimes occurs. [See Burck in Ann. Buitenz. 
III. 1883, and iv. 1884.] 
Several R. are of economic importance, e.g. Cinchona, Coffea, 
Uragoga, Rubia, &c. 
Classification and chief genera (after Schumann). The R. are 
closely allied to Caprifoliaceae (q.v.) and less nearly to Compositae &c. 
I. CINCHONOIDEAE (ovules 00 in each loculus). 
A. Cinchoninae (fruit dry) : 
a. Firs, solitary or in decussate panicles. 
a. Fir. regular; seed not winged; corolla valvate. 
1. Condamineeae: Condaminea. 
2. Oldenlandieae : Oldenlandia, Houstonia, Pentas. 
b. As a, but corolla imbricate or convolute. 
3. Rondeletieae : Rondeletia. 
c. As b, but seed winged. 
4. Cinchoneae: Cinchona, Bouvardia, Cosmibuena. 
d. As a, but corolla 2-lipped. 
5. Henriquezieae : Henriquezia. 
/ 3 . Firs, in heads. 
6. Naucleeae : Uncaria, Nauclea. 
B. Gardeninae (fruit fleshy) : 
7. Mussaendeae (corolla valvate) : Mussaenda. 
8. Gardenieae (corolla imbricate or convolute) : Randia, Gar- 
denia, Posoqueria, Duroia. 
II. COFFEOIDEAE (ovules 1 in each loculus). 
A. Guettardinae (ovule pendulous; micropyle facing upwards): 
9. Albert eae: Alberta. 
10. Knoxieae: Knoxia. 
11. Vanguerieae: Plectronia, Cuviera. 
12. Giiettardeae : Guettarda. 
13. Chiococceae : Chiococca. 
B. Psychotriinae (ovule ascending; micropyle facing down- 
wards) : 
a. Corolla convolute. 
14. Ixoreae : Coffea, Ixora, Pavetta. 
p. Corolla valvate. 
a. Ovules inserted at base of loculus. 
15. Psychotricae: Psychotria, Rudgea, Uragoga, Lasianthus, Myr 
mecodia. 
16. Paederieae: Paederia. 
17. Anthospermeae : Nertera, Coprosma, Mitchella. 
18. Coussareeae: Faramea. 
