312 
A MEMOIR UPON THE GENUS 
as in the larger species. The subdivisions of the proximal face of the cuboid in P. 
'paludosus are not well marked. The calcaneal porticm is very deep and slightly 
convex from side to side ; the astragalar facet is elongate and narrow, larger above 
than below and concave from above downward. The facets for the navicular and 
ectocuneiform are short and deep ; they are qxiite different in form from those of the 
smaller species. The cuboid-ectocnneiform facet is much larger than the facet for 
the navicular. The distal face of the cuboid in P. pahidostis is much deeper than 
broad ; its posterior portion is narrower than the anterior. The surface of this facet 
is plane and slightly oblicpie. The inferior internal angle of this cuboid exhibits a 
small truncated facet for metatarsal III. Another cuboid in the collection does 
not show any facet for this metapodial (No. 10,282). 
Metatarsals . — The metatarsal region of P. paludos 2 is is short, with the digits 
wide-spreading. The metatarsal 111 is slightly larger and broader than the lateral 
metapodials, but does not show the difference in size that is seen in the foot of Palce- 
othermm.^ for example. The proximal portion of metatarsal II is broad, with 
the external side abruptly truncated. A peculiarity of this metatarsal is that 
it shows no facet for the entocuneiform such as occurs in Limnohyops. Its 
articular face for the ectocuneiform exhibits two round separated facets. The distal 
extremity of this metatarsal is heavy, with prominent tuberosities Ijordering the 
articular surfaces.- The upper part of metatarsal III is very narrow compared with 
its distal portion. The surface for the ectocuneiform is nearly flat and obliquely 
placed ; its cuboidal sides show two facets for metatarsal IV. The distal part of 
metatarsal III is extremely heavy and its articular surface is wide and subdivided 
posteriorly by a prominent keel. 
Metatarsal IV is strongly concave on its external side and its shaft is much 
heavier than that of metatarsal II ; its superior face is unusually Hat, being 
bread and shallow. The metatarsals in P. pahidosus resemble closely those of the 
smaller species of the genus, the difference being largely in their heaviness and the 
size of their articulating surfaces. 
Summary or the Characters and Affinities of the Skeleton of Pal.eosvops 
PALUDOSUS. 
In the foregoing chapters I have described the bones of the skeleton of this 
animal and compared them with those of recent and fossil Perissodactyles, and 1 
now propose, as briefly as possible, to sum up the ^irincipal characters of the skeleton, 
and to compare its peculiarities with those of allied forms. 
Certain characters are found in the vertebral column which depart widely 
from those of the Tapir. These are the form and position of the articular pro- 
cesses of the posterior axial region. It was also observed that among recent forms 
the Horse ajiproaches PalcBosyops more nearly than any other existing Perissodac- 
tyle in the position of its lumbar processes. 
