486 THE STRUCTURE AND RELATIONSHIPS OF ANCODUS. 
with it, and of tlie tarsus only the calcaneum and astragalus. In a second fragmen- 
tary skeleton the metatarsals and metacarpals are of nearly the same length, while 
a third specimen, a beautifully preserved hind-foot, of which the calcaneum and 
astragalus agree closely in size with those of the first-named specimen, the metatar- 
sals much exceed in length the metacarpals of the latter. A more extensive series 
of fore- and hind-feet associated together will be required to determine accurately 
these relations. In the following description the isolated pes is made use of. 
The metatarsals differ from those of Oreodon in the greater proportionate de- 
velopment of the median pair and reduction of the laterals. The first metatarsal 
is a small nodular rudiment of irregular shape, which is broadest proximally, taper- 
ing distally to a blunt point, and is of roughly trihedral section. It is attached to 
the distal end of the entocuneiform, but there is no clearly defined facet, and doubt- 
less the joint was very imperfectly developed. No phalanges are connected with 
the rudimentary metatarsal, and the hallux is thus far more reduced than is the 
pollex, which is relatively larger than in the dogs. The rudiment is entirely plantar 
in position, being concealed from the front by mt. II, behind which it lies. 
The second metatarsal is long, much compressed laterally, and of trihedral 
shape, expanded and somewhat thickened distally. The head is very narrow ; it 
bears a small facet for the mesocuneiform, and abuts by a minute surface against 
the tibial side of the ectocuneiform, beneath which the shaft is excavated to receive 
an expansion of the head of mt. III. The postero-internal angle is cut away at the 
proximal end, forming a long, oblique surface, overlapped by the wide entocuneiform. 
The distal epiphysis has been lost from the specimen. 
The third metatarsal is very long and, though much heavier in every way than 
the laterals, is slightly shorter and considerably narrower than mt. IV, while in 
Oreodon it is somewhat longer. The proximal end rises less above the head of 
mt. IV, and has a smaller contact with the cuboid than in the specimen figured by 
Kowalevsky (PI. XXXVIII, fig. 1) more so than in that figured by Filhol (PI. 25, 
fig. 124), and the head is relatively narrower than in either. On the fibular side, 
below the head, is a deep pit into which is received a corresponding projection 
from mt. IV. By means of this and the mode of articulation with the tarsals 
the median pair of metatarsals are very firmly locked together. From the plantar 
side of the head arises a long and stout projection, which is held in place by the 
cuboid proximally, the entocuneiform' internally, and mt. IV externally The 
shaft IS more compressed antero-posteriorly, and has a more flattened dorsal face 
than m OWOB. The distal trochlea is wider proportionately than in that genus, 
but IS demarcated from the shaft in the same way by a deep pit on the dorsal face 
of ‘he bone. In all the metatarsals the oarlna is entirely plantar, 
not mneh e“ve d ‘'l' <>f i <* 0 ^“ 
the proxi,:aTe„d.” B™ K “t’sSl d 
the European species it is rather mt rnder^r^f^I^I. 
are exclnswely w.th the cuWd, but are nevertheless quite 00^^ On Z 
