CONTRIBUTIONS TO BOTANY. 
257 
of the cell, where the ovules became ensconced ; the parietal 
emanations tapered upwards, gradually closing and joining 
together so as to fomi the style and stigma ; and while the cells 
thus produced continue to grow downwards, the anatropous 
ovules remain attached to their original placentary supports, 
with their micropyles pointing upwards. At this stage, 31. 
Baillon’s description ceases, and he does not relate what occurs 
in the fmther development of Cordia^ or what takes place in 
Heliotropuim or Ehretia. The differences in these subsequent 
developments may, however, be summarized in the following 
manner. 
In Gordia the four parietal emanations gradually approach 
each other, to form four equal partitions united in the axis, so 
that, at the period of flowering, the ovaiy is completely 4-celled, 
with a single subanatropous ovule in each cell, attached to the 
internal angle by a point below its apex or by its middle : the 
com])letlon of this growth results in a drupaceous fruit, with a 
4-celled osseous nut, each cell with a single seed suspended 
from below the summit, with a descending raphe terminating 
in the basal chalaza, the radicle of the exalbuminous seed 
being superior, and the cotyledons singularly plicated length- 
wise. In the base and centre of the nut a large hollow is seen 
filled with placentary tissue, from Avhich four sets of nourish- 
ing vessels issue, penetrating through a minute perforation 
near the summit of each cell, and terminating in the hilar 
attachment of the seeds. These are the very peculiar trans- 
formations that serve to distinguish the Cordiacece from all 
other developments of the same alliance. 
In Ehretia^ the ovary at an early stage is developed much 
after the manner of Gordia : the rudiments of four ovules 
emanate at the same points, and we see four similar inter- 
mediate parietal enlargements ; but the basal placentary ridges 
combine to form a compressed elevated line, ranning from 
front to back across the axis, which continues to grow upwards, 
carrying the ovules with it, or, what amounts to the same tiring, 
the principal growth of the whole takes place downwards ; and 
in this manner the placentary columella is produced, whicli 
31. Baillon does not seem to have noticed. On the other hand, 
the parietal enlargements do not meet round the axis, as in 
Gordia^ but are thrust aside after a while ; the sinister and 
dexter emanations form semisepta, which, on approaching the 
columella, become suddenly reflected both ways, in parallel 
directions, to meet the corresponding parietal growths from the 
anterior and posterior walls : the consequence is that, at the 
period of the perfection of the flower, we see two bilocular 
carpels, each cell having a suspended ovule, while a vacant 
2l2 
