221 
whole of the branch surfaces are eelluliferous without any separation by ridges. The 
separation of the two genera is not a question of which aspect of the polyzoarium is 
cell-bearing, but of the arrangement of those cells on the branches. 
It is, howerer, to Poh^pora, as pointed out by Dr. Waagen, as well as to Phyllopora, 
that Protorefepora is most nearly allied. Here the arrangement of the cells on the 
interstices is identical, the only question being as to which aspect of the polyzoarium is 
eelluliferous. In Protoretepora it is undoubtedly the inner. I quite agree with Dr. 
Waagen as to the close affinity of Poh/pora and Protoretepora ; but before they can be 
United it must be shown that in the type species on which McCoy established the former, 
the polyzoarium is eelluliferous on the inner face, and in consequence therefore infundi- 
buliform. Until this is demonstrated Protoretepora can be satisfactorily made use of as 
u genus. I cannot agree with Dr. Waagen that the question of this, or that face being 
Cell-bearing, is one of accident. Tie says — “ The sole difference between the two consist- 
i^ug in the circumstance, that in the one the pores are said to be placed on the inner, in 
e other on the outer side of the funnel-shaped colony. Such a position of the pores is 
evidently only accidental, depending upon the mode in which the quite young and still 
an-shaped colony first became twisted ; if towards the poriferous side the pores remained 
on the inner side, if in the reverse direction the pores remained outside.”* That “the 
position of the pores is evidently accidental ” is pure assumption, and it has yet to be 
proved that these infundibuliform genera commenced life as a “still fan-shaped colony,” 
uud not with a miniature infundibuliform or cup-shajied outline. I much question if the 
uiode of growth is not a regular and constant factor in the anatomical construction of 
such genera. It certainly is constant in a large number of recent Polyzoa, examined 
the purpose by Mr. T. Whitclegge, of the Australian Museum, and the Writer, 
'■ing Petepora as an example of the more or less infundibuliform solitary polyzoaria, 
found that the inner aspect was invariably eelluliferous in both B. monilifera, 
'Ificq the Port Phillip form, and Jl. jacksoniensis, Busk., inhabiting Port Jackson, 
nd the outer side of the polyzoarium never. The same is the case in such tube-like 
species as Mucronella delicatula, Busk., and M. Usinuala, Smith, from Thursday 
d also Scliizoporella australis, Tlasw'". The fact is again repeated in the sub- 
endroid species of Gellepora, and in saucer-shaped polyzoaria, like Carlasea crihiformis. 
^ One very essential character ascribed by De Koninck to his genus appears to have 
• overlooked by Dr. W aagen. The former says of Protoretepora, “ toute la surface 
^ erne est ornee de plusiours lignes de petites cellules.” Now, whatever the eelluliferous 
spot may have to do with generic separation, we have here a feature, which on Dr. 
aagen’s own showing is of importance, for, he says, speaking of Polypora, “the 
1 ores restricted to the branches.” Until, therefore, McCoy’s typo species of Polypora 
ain P eelluliferous on the inner aspect, and in the case of both Phyllopora 
ror • ^^t/pora on the dissepiments as well as the interstices, De Koninek’s genus can 
ain its rank as such. 
^ Protoeeteporx ampla, Lonsdale, sp. 
ampla, Lonsdale in Darwiu’.s Geol. Ob.s. Vole. Island.s, 1844, p. IfiS. 
>. Lonsdale in Strzelecki’s Phys. Deserip, N. S. Wales, &c., 1845, p. 268, t. 9, if. Za-d. 
^olvn ” Wilkes’ IT. S. Explor. Exped., 1849, p. 710, t. 11, If. 1 and la. 
^rot'^ Etheridge fil., Trans., R. Soc. Viet., 1876, xii., p. 66, Pi., f. 1. 
°retcpora ampla Iv^rs), Etheridge fit. Cat. Australian Eoas., 1878, p. 45. 
less Ctliar. Polyzoarium funnel or cup-shaped, with the upper portions more or 
dis '^5^i®pled. Interstices broad, flattened, and expanding previous to bifurcation ; 
^__^*®ents short, sometimes eelluliferous. Uenestrules oval, and rather long, the 
* Pal. Indioa {Salt Range Fos,?.), 1885, Ser. xiii., No. 1, fasc. 5, p. 775. 
