14 
Indiana University Studies 
Figs. 32 and 33. Fig. 32. Dorsal and ventral views of the fruits 
of Polytoca. Fig. 33. Dorsal and ventral views of the fruits of 
Sclerachne. In each group the caryopsis is shown in the figures in the 
middle. 
Figs. 34-36. Pairing of spikelets in Sclerachne. S, staminate 
spikelet; F, pistillate spikelet; R, rachis; St, sterile pedicel accompany- 
ing pistillate spikelet; P, pedicelled staminate spikelet; S, sessile stami- 
nate spikelet. The spikelet terminating the rachis is sessile, and its 
pedicelled mate is lacking. 
is consistently taller and more slender. The leaves and stems 
are glabrous or somewhat ciliate. 
The homologue of the spathe seen in Polytoca is here more 
of the nature of a foliage leaf. The spike bears one or two 
basal pistillate spikelets each, accompanied by a rudiment, and 
above these only one to three staminate spikelets (Fig. 23). 
Altho modified by the small number of staminate units, the ar- 
rangement of the spikelets is the same as in Polytoca (Figs. 
34-36). 
No significant peculiarity of the male or female spikelets 
has been noted, except that in the former the functional 
androecium is sometimes reduced to a single stamen. The 
ligule-like appendage at the top of the outer glume of the 
