IOWA ACADEMY OP SCIENCE 
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Lamarckiana. The leaves in the upper part of the stem in 0. Lamart- 
kiana, are, however, frequently nearly or quite smooth. 
AA^e have shown that the first Oenothera introduced into Europe from 
Virginia was more closely related to the 0. Lamar ckiana of our present 
cultures than to any other form, differing from it only in such minor 
points as the width of the rosette leaves ; which seem also to have been 
of a iialer green color, because the Bauhin description and Parkinson 
agree on this point. But the important independent description of 
Hernandez in 1651 definitely refers to the crinkling of the leaves. This 
makes it highly probable that the plant of Hernandez was almost or 
quite identical with our 0. Ij am, ar ckiana, Ser. 
O. GRxVNDIFLORA AIT. 
The history of the discovery of 0. grandiflora in Alabama and its in- 
troduction into Kew has already been given by DeVries (1901) and par- 
ticularly MacDougal (1905, p. 7), and need not be repeated here. But 
certain interesting data can be added. Before entering upon these it 
will be advantageous to outline some of the differences between 0. Lam- 
ar ckiana and 0. grandiflora as we now know them from cultures (see 
Vail, 1907, p. 66; Grates, 1909c, p. 131). In 0. grandiflora the buds bear 
only a short and inconspicuous type of hair, giving them an almost gla- 
brous appearance (in some cases entirely glabrous), while in 0. Ijam- 
arckiana and all its mutants there is in addition a long, curved type of 
hair, arising from papillae and giving the buds a pubescent appearance. 
The same is true of 0. hiennis. The same type of hair is found on the 
stems in 0. Lamar ckiana and 0. hiennis, arising from papillae which are 
usually red, so that the stem is covered with small red dots. 0. grandi- 
flora from Alabama shows the same condition on the stem, but in some 
of the forms of 0. grandiflora from near Liverpool, England, the long 
type of hair is frequently almost wholly absent, leaving the stem nearly 
glabrous. The buds in 0. grandiflora are also more slender and rounded, 
and the sepal tips frequently longer and usually more setaceous than in 
0. Lamar ckiana. 
In addition the rosettes are very unlike, the leaves in 0. grandiflora 
being smooth, thin, and with a series of characteristic basal lobes, while 
in 0. Lamarckiana they are crinkled, thicker, and without the basal lobes. 
But unfortunately, the rosettes are rarely mentioned, except in connec- 
tion with recent studies and cultures, and in the very early works. 
Professor DeVries has given an account of the history and synonomy 
of 0 . grandiflora Ait. He prefers to call it 0 . snaveolens, the name in- 
troduced by Desfontaines, to avoid confusion on account of the various 
