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Indiana University Studies 
Cynips subgenus Atrusca, new subgenus 
agamic forms 
Dryophanta Mayr, 1881 (in part), Gen. gallenbew. Cynip.: 35. Mayr, 
1886, Verh. zoo.-bot. Ges. Wien 36: 370-371. Dalla Torre, 1893 (in 
small part), Cat. Hymen. 2:48. Dalla Torre and Kieffer, 1902 
(in part), Gen. Ins. Hymen. Cynip.: 52. Beutenmiiller, 1911 (in 
part), Bull. Amer. Mus. Nat. Hist. 30: 343-352. Also of other 
authors. 
Loxaulis of one assignment: Ashmead, 1885, Trans. Amer. Ent. Soc. 
12: 296. 
Diplolepis Dalla Torre and Kieffer, 1910 (in part), Das Tierreich 24: 
342. Weld, 1926 (in small part), Proc. U.S. Nat. Mus. 68 (10): 
18-23. Also of other assignments and other authors. 
Cynips of some assignments of early authors. 
Holcaspis of some assignments of authors. 
Disholcaspis of several assignments of authors. 
Andricus of two assignments (Ashmead, Kinsey). 
FEMALE. — The cheeks hardly at all enlarged behind the eyes; 
antennae of moderate length, shorter in all forms with shortened wings; 
with (rarely) 13 or (usually) 14 segments; the thorax of moderate 
size, smaller in shorter-winged varieties; parapsidal grooves continuous; 
median groove lacking; mesopleuron entirely punctate; abdomen entirely 
smooth and naked except for the hairs latero-basally; hypopygial spine 
rather long, slender, bluntly pointed, somewhat but not greatly widened 
well back from the tip; tarsal claws rather slender, broadened basally 
but only weakly toothed (more distinctly toothed in centricola) ; wings 
long, about 1.35 times the body in length; or wings shortened to 1.15, 
0.90, or 0.85 times the body in length; if the wings are long, the second 
abscissa of the radius is well curved for its entire length (less curved 
in centricola ), more strongly curved terminally, the tip of the vein 
angulated and swollen, the radial cell distinctly short and broad (longer 
in centricola) and with or without spots, the areolet very small to 
large, the cubital cell with a clouded patch basally and numerous spots 
apically, and the discoidal cell with or without patches or spots; in 
shorter-winged varieties the venation is reduced but all the veins are 
represented; length 2.0 to 4.5 mm., averaging rather large but not stout 
(except in centricola). 
GALL. — Large, strictly spherical unless drawn out basally or ab- 
normally distorted; entirely smooth and naked unless for a puberulence 
or microscopic pubescence; the outer walls of the gall thin, soft and 
pliable when moist, hard and brittle when dry, bright rose red and 
green when young, with or without spots, becoming light rose brown 
when older; the gall entirely hollow except for the thick-walled, cen- 
trally placed larval cell and for a moderate number of fine, silky, radiat- 
ing fibers that connect the larval cell and the outer walls of the gall. 
