212—4 INDIAN TERTIARY AND POST-TERTIARY VERTEBRATA. 
tertiary of the Maltese Islands by T. champsoides (Owen), and T. gaudense (Hulke). 
The former differs from T. schlegeli by the presence of an additional premaxillary 
tooth, and thereby approaches Gharialis gangeticus. A crocodilian from the (ap- 
parently) miocene of Eggenburg in Lower Austria, described by Messrs. Toula and 
Kaih under the provisional name of Gharialosuchus^ eggenhurgensis^ has likewise been 
shown by the present writer in the paper quoted to be inseparable from Tomistoma, 
and may be known as 2\ eggenhurgense. In this species the anterior border of the 
orbit is prominent, as, in Gharialis gangeticus^ and there is likewise an additional tooth 
in the premaxillse, between the proper first and second teeth : it is possible that it 
may be specifically the same as T. champsoides, in which case the former name has 
the right of priority. A new gharialoid genus from the Siwaliks is described in the 
sequel, and forms the second division of the gharialoid group in the table given 
above. 
Genus I. : CROCODILUS, Linn.® 
Definition. — It may perhaps be doubted if Crocodilus toliapicus, Owen,^ from the 
London Clay, can really be included in this genus, as it has ^ teeth, and the mandi- 
bular symphysis is elongated (although not to the same extent as in Tomistoma 
schlegeli) ; it agrees, however, with typical species of Crocodilus (and thereby differs 
from Tomistoma) by the extension of the nasals to the anterior narial aperture^ 
Crocodilus champsoides, Owen,® from the same formation, has 21 upper teeth, but agrees 
with C. cataphractus (in which the teeth number ff) in the expansion of the premaxillse, 
and their perforation by the fourth mandibular tooth, as well as in the relation of 
the nasals to the external nares. Including the latter species, the genus may be 
provisionally defined from cranial characters, as follows : — Snout moderately long, or 
short, nasals articulating with premaxillse,® anterior border of orbit not prominent, 
third and ninth upper and fourth lower teeth generally larger than the others, first 
mandibular tooth received into a pit or perforation, fourth into a lateral notch in the 
cranium. The muzzle is longest, and the teeth are less differentiated in the gharialoid 
forms like C. cataphractus, and G. champsoides, which not improbably indicate a 
transition towards Thoracosaurus and Tomistoma. 
Distribution. — At the present day the genus occurs in America, Africa, Asia, and 
Australia ; it is strongly developed in nearly all the tertiaries, being represented in 
the London Clay’’ of England by the long-snouted C. champsoides, and in the middle 
eocene by the short-shouted C. hastingsice, Owen.® 
1 ‘Anzeig. k Ak. Wiss.’ 1885. pp. 107-109. 2 Amended from Gavialosuchus. 
3 ‘ Syst. Nat.’ ed. 12. vol. I. p. 359 (1766). As a subgenus of Laccrta. 
4 ‘ Fossil Eeptilia of London Clay, etc.’ pt. 2. p. 29. pis. II. IIA. (1850). 5 Hid. p. 31. pi. III. 
6 In some instances (e.y. C. hastingsice) the nasals do not reach the anterior nares, although they do so in most species. 
7 0. icenicus and C. cantabrigiensis, Seeley (Quart. Journ. Geol. Soc.’ vol. XXX. p. 693 [1874], and XXXII. p. 437 
[1876]), being founded on vertebrae, their generic determination is uncertain. 
5 Op. cit. p. 37. pis. VI. VII. 
