SIWALIK CKOCODILIA, LACERTILIA, AND OPHIDIA. 
19—227 
indicating the affinity of the species to Oharialis. The dental alveoli (fig. 4a.) are 
separated from one another by an interval somewhat less than their antero-posterior 
diameter ; and between each alveolus there is a broad, shallow, pit for the reception 
of the tip of a mandibular tooth. In the presence of these interdental pits the 
specimen differs from Oharialis gangeticus ; while in their position it agrees with 
Tomistoma schlegeli, although the depth of the pits themselves is very considerably 
less. 
Distinctness and afflnities . — That the present form is distinct from each of the 
three species already described is sufficiently apparent from the description ; and it 
is equally distinct from the so-called G. niacrorhgnchus, and also from G. dixoni. The 
relations of the nasals indicate that it should be referred to the genus Oharialis, 
although the interdental pits of the maxillae show an approximation to Tomistoma. 
It is much to be desired that the cranium proper and the termination of the rostrum 
of this interesting form may one day be forthcoming, when its true affinities may be 
more fully indicated. 
Distribution. — All the known remains of G. leptodus have been obtained from 
the true Siwalik Hills and the eastern Punjab. 
Species 5. GtHaeialis pachyehynchus, n. sp. nobis. 
History. — The type specimen of this species is described for the first time ; a 
second specimen has, however, been noticed by FalconeP under the name of 
Leptorhynchus. 
Cranial rostrum. — The specimen of which three views are given in plate XXXIII. 
figs. 1, la, lb, was collected by Mr. F. Fedden in the lower Siwaliks of the Laki 
Hills, Sind, and indicates a species of gigantic size. It comprises the extremity of 
the cranial rostrum, extending as far back as the fourth tooth behind the notch for 
the fourth mandibular tooth. On the palatal aspect (fig. la) it will be seen that 
while some of the teeth, although broken, still remain in their alveoli, others have 
fallen out, and their alveoli are filled with matrix. As the general contour of the 
specimen is essentially gharialoid, comparisons may be restricted to that group. The 
termination of the premaxillm much resembles that of the existing gharial, containing 
one large tooth on either side, the outer border of which is bounded by a deep 
notch ; while the extremity of the premaxillse is strongly deflected. The cavity for 
the reception of the first mandibular tooth does not extend on to the facial aspect of 
the rostrum (fig. 1.), but is roofed over with bone, and would only require the com- 
pletion of the outer wall to form it into a pit ; as, however, the first mandibular 
tooth would be entirely visible externally if the mouth were closed, the cavity in 
question may still be termed a notch. The second^ and third teeth are of very large 
size, but the fourth appears to be absent ; the widest part of the premaxillae is at the 
1 “Catalogue of Fossil Vertetrata of Asiattc Society of Bengal,” p. 258, No. 14 (1859). 
2 A depression on the posterior side of the notch for first mandibular tooth appareatly indicates the alveolus of an 
accessory tooth between the proper first and second upper teeth. 
