234—26 INDIAN TERTIARY AND POST-TERTIARY VERTEBRATA. 
give it a distinctly pitted appearance : tlie same feature is apparent in the mandible 
of the type specimen. 
Dimensions . — The following dimensions taken from the specimens described 
above are compared with those of an adult skull of Qharialis gangeticus measuring 
thirty-one inches from the occipital condyle to the extremity of the premaxillse. 
G. gangeticus. 
Fossil. 
Width of cranium in advance of palatal foramina 
4-8 
120 
,, ,, at 7th tooth from do. 
3-1 
10-4 
,, mandible at fourth tooth in advance of hinder border of symphysis. 
2-9 
9-6 
Length of rostrum to 1 6th tooth 
15-4 
27-0 
Width of articular surface of quadrate and quadrate -jugal .... 
2-3 
6-2 
These dimensions indicate an animal of about three times the size of the specimen of 
G. gangeticus., but with a relatively shorter rostrum. Assuming that the cranium of 
the existing species belonged to an individual close on twenty feet in length, the 
fossil species would have attained a length of between fifty and sixty feet. 
The centrum of an early dorsal vertebra from the Siwalik Hills in the British 
Museum may, from its gigantic size, be pretty safely referred to the present form. 
The vertical diameter of the anterior articular cup of this specimen is 4‘4, and the 
transverse 3 ‘9 inches : and it indicates an individual fully three times the size of the 
largest recent crocodile skeleton in the British Museum. A dorsal scute in the same 
collection (No. 17066) from the Siwalik Hills has a transverse diameter of 7*1, and 
an anterior-posterior of 4*2 inches. 
Teeth . — In the subequal size of the maxillary and the majority of the hinder 
mandibular teeth the present form differs from the true crocodiles and alligators, and 
agrees with Gharialis and Tomistonia ; and it appears that in proportion to the size of 
the rostrum the teeth are not larger than in Tomistonia champsoides of the miocene of 
Malta.^ In the upper teeth the fore-and-aft ridges are placed more mesially than in 
the fourth upper tooth of Grocodilus, and their inner surfaces are less flattened. 
Instead of the ridges being directed nearly laterally as in G. gangeticus, they are 
placed obliquely as in T. schlegeli. The Indian Museum possesses numerous teeth 
from the Siwaliks of the Punjab which apparently agree with those of the present 
form. The specimen represented in plate XXXIII. fig. 5 closely resembles one of 
the teeth of the anterior third of the type mandible. 
Distinctness and affinities . — The foregoing comparisons decisively indicate that 
the affinities of the present form are with the gharialoid group. That it is distinct 
from Gharialis 'pachyrhynchus {ffiisXQ XXXIII. figs. 1, 2) is evident from the absence 
of the premaxillary expansion, by the presence of the pit for the fourth mandibular 
tooth, the relations of the premaxillary to the maxillary dental series, and the 
apparent absence in the latter of the alligatoroid relations of the upper and lower 
dental series. It is equally evident that it is specifically distinct from all the other 
Siwalik forms already described. 
With regard to generic characters, the present form differs from both Tomistonia 
1 See a paper by the present writer in the ‘ Quart. Journ. Geol. Soc.’ vol. XLII. (1886). 
