THE BRACHYURA AND MACRURA OF PORTO RICO 
29 
Eupanopeus occidentalis (Saussure). 
Panopeus occidentalis Saussure, Rev. Mag. Zool. (2), ix, 502, 1857; Benedict & Rath bun, Proe. U. S. Nat. Mus., xiv, 3fi0, pi. xx, 
f. 3, pi. XXIII, f. 14, 1891. 
Eupanopeus occidentalis Rathbun, Bull. Lab. Nat. Hist. State Univ. Iowa, iv, 273, 1898. 
Differs from E. herbstii as follows: Gastric region more elevated, independently convex. Second 
tooth of lateral margin narrower and separated by a deeper sinus from first tooth; third to fifth teeth 
thicker, more prominent and more widely separated from each other, third tooth with anterior and 
exterior margins subequal. Front more advanced. Carpus of chelipeds with a groove parallel to distal 
margin. Dark color on thumb not continued on palm. Ambulatory legs a little longer and more 
slender. Abdomen of male wider, especially noticeable at penult segment. 
In alcohol the chelipeds have a pinkish tinge not seen in E. herbstii. 
Male: Length, 20 mm. ; width, 27.2 mm.; fronto-orbital width, 16 mm.; width of front, 8.4 mm. 
South Carolina to Pernambuco; Bermudas. Hucares, Porto Rico, 2 females, 1 with Sacculina. 
Eupanopeus americanus (Saussure). 
Panopeus americanus Saussure, Rev. Mag. Zool. (2), ix, 502, 1857; M6m. Soe. Phys. Geneve, xiv, 432, pi. I, f. 8, 1857. 
Panopeus meolatus Benedict & Rathbun, Proc. U. S. Nat. Mus., xiv, 361, pi. xxi, f. 3, 1891. 
Eupanopeus americanus Rathbun, Bull. Lab. Nat. Hist. State Univ. Iowa, iv, 273, 1898. 
The carapace is narrower (width 1.3 times length) and flatter than in E. herbstii. The transverse 
rug* are numerous. Sinus between first and second lateral teeth deepish; other teeth broad, rather 
blunt, third tetragonal, fourth with anterior margin at right angles to median line. Lobes of front with 
more oblique margins than in E. herbstii. Chelipeds similar to those of E. herbstii; a faint carpal groove 
may be present. Tooth at base of dactyl longer than in E. herbstii. Abdomen of male narrow, as in 
E. herbstii, but in shape resembling that of E. occidentalis. 
Male: Length, 19.5 mm. ; width, 26.3 mm. ; fronto-orbital width, 15.5 mm. ; width of front, 7.5 mm. 
Bahamas to Brazil. Arroyo, Porto Rico; 1 male. 
Eupanopeus bermudensis (Benedict A Rathbun). 
Panopeus wurdcmamiii Benedict & Rathbun, Proc. U. S. N. M., xiv, 372, pi. xxiv, f. 6 and 7, 1891 (not P. wurdemannii Gibbes). 
Panopeus bermudensis Benedict A Rathbun, Proc. U. S. N. M., xiv, 376, pi. xx, f. 2, pi. xxiv, figs. 14 and 15, 1891. 
Eupanopeus bermudensis Rathbun, Bull. Lab. Nat. Hist. State Univ. Iowa, iv, 273, 1898. 
A smaller species than any other Eupanopeus taken at Porto Rico. The carapace more deeply 
areolated than any of above; a well-marked transverse line on branchial region on a level with last 
lateral sinus; also several shorter granulate ridges, and granules toward front and antero-lateral margins; 
otherwise smooth. Second tooth of lateral margin lobiform, separated from first by a rather shallow 
sinus; third and fourth large, with arcuate outer margins, third obtuse, fourth subacute; fifth narrow 
and pointed. The front has a thick, obliquely beveled edge, obscurely grooved, deflexed; in a dorsal 
view the lobes are nearly straight, oblique, but in a front view very sinuous. The carpus of cheli] eds 
has a well-marked groove; fingers strongly bent down and usually light horn-colored with white tips; 
the color extends partially on palm also. Legs slender and delicate. Third segment of abdomen of 
male narrow, with rounding posterior corners, not reaching cox* of fifth pair of feet. 
Length of male, 5.8 mm.; width, 8 mm. ; fronto-orbital width, 5.5 mm.; width of front, 2.8 mm. 
Bermudas; Florida Keys to Brazil. Porto Rico: Mayaguez; Boqueron Bay; GuanieaBay; Arroyo, 
on Lighthouse Reef; Ensenada Honda, Culebra. Also taken at Nassau, Bahamas; 1 specimen having 
the genital appendages of a male, and an abdomen with seven segments, as in an immature female. 
Eupanopeus harttii (Smith). 
Panopeus harttii Smith, Proc. Boston Soc. Nat. Hist., xii, 280, 1869; Trans. Conn. Acad. Sci., li, pp. 5 and 34, pi. i, f. 5, 1869. 
Eupanopeus harttii Rathbun, Bull. Lab. Nat. Hist. State Univ. Iowa, iv, 273, 1898. 
Carapace little convex, rugose, regions very distinctly marked. First and second antero-lateral 
teeth small, tuberculiform, separated by a U-shaped sinus. Remaining teeth rather narrow, thickened; 
tips tuberculiform. Front little advanced and with a small but distinct tooth at the orbital end. 
Carpus of chelipeds rugose and with a deep anterior groove. Color of lingers not continued to the 
palm. Abdomen of male reaching coxal joints. 
Length of male, 8.6 mm.; width, 12.6 mm.; fronto-orbital width, 8.6 mm.; width of front, 4.1 
mm. A small species, but larger than E. bermudensis. 
Florida Keys to Brazil. Porto Rico: Boqueron; Mayaguez; Mayaguez Harbor. 
