20 
C. D. Camp, C. W. Condee, and D. G. Lovell 
in Alabama (stage 21). Meeks and Nagel (1977) reported a similar 
hatching size to that in this study but did not report the larval stage at 
hatching. Herreid and Kinney (1967) reported that Alaskan wood frogs 
hatch at larval stage 20. Tadpole development time in our study was 
longer than that reported for wood frogs elsewhere (Hinckley 1882, 
Beilis 1957, Herreid and Kinney 1967, Meeks and Nagel 1977, Berven 
1982b, Davis and Folkerts 1986). Berven (1982b) suggested that temper- 
ature was a major factor in variation in wood frog larval development 
periods. 
Maximum total length reached by tadpoles in Georgia was similar 
to that reported from most parts of the species range (Beilis 1957, 
Herreid and Kinney 1967, Meeks and Nagel 1977). Size at metamorphosis 
was similar in Georgia wood frogs to that reported elsewhere (Hildebrand 
1949, Beilis 1961, Meeks and Nagel 1977, Berven 1982b). 
Catastrophic mortality as a result of premature desiccation of NTL 
and RC1 emphasizes the risky nature of breeding in temporary pools. 
Desiccation has been identified as a major selective force for amphibians 
breeding in temporary ponds (Semlitsch 1987). 
ACKNOWLEDGMENTS.— We thank Carlton Bowers for access to 
wood frog breeding sites on his property and Trip Lamb, Ray Semlitsch, 
and J. Whitfield Gibbons for their comments on the manuscript. 
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LeConte. Ph.D. dissertation, Univ. Minnesota, Minneapolis. 
Beilis, E. D. 1961. Growth of the wood frog, Rana sylvatica. Copeia 
1961:74-77. 
Berven, K. A. 1982a. The genetic basis of altitudinal variation in the wood 
frog Rana sylvatica. I. An experimental analysis of life history traits. 
Evolution 36:962-983. 
Berven, K. A. 1982b. The genetic basis of altitudinal variation in the wood frog 
Rana sylvatica. II. An experimental analysis of larval development. Oecologia 
52:360-369. 
Davis, M. S., and G. W. Folkerts. 1986. Life history of the wood frog, Rana 
sylvatica LeConte (Amphibia: Ranidae), in Alabama. Brimleyana 12:29-50. 
Gibbons, J. W., and R. D. Semlitsch. 1982. Terrestrial drift fences with pitfall 
traps: an effective technique for quantitative sampling of animal populations. 
Brimleyana 7: 1-16. 
Gosner, K. L. 1960. A simplified table for staging anuran embryos and larvae 
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