Invertebrate Cave Fauna 
15 
Table 1. Frequency distribution by order or subclass of troglobites and 
troglophiles in the study area. 
Order or Subclass 1 
No. of Troglobites 2 
No. of Troglophiles 2 
Alloeocoela (flatworms) 
1 
_ 
Tricladida (flatworms) 
4 
2 
Lumbriculida (oligochaetes) 
3 
- 
Mesogastropoda (snails) 
2 
2 
Stylommatophora (snails) 
1 
2 
Amphipoda (amphipods) 
20 
1 
Isopoda (isopods) 
15 
3 
Decapoda (crayfishes) 
Pseudoscorpiones 
- 
1 
(pseudoscorpions) 
15 
2 
Acari (mites) 
2 
3 
Opiliones (harvestmen) 
- 
2 
Araneae (spiders) 
8 
9 
Lithobiomorpha (centipedes) 
1 
- 
Spirostreptida (millipeds) 
- 
1 
Chordeumatida (millipeds) 
9 
7 
Julida (millipeds) 
- 
1 
Collembola (springtails) 
4 
8 
Diplura (bristletails) 
6 
- 
Orthoptera (crickets) 
- 
1 
Coleoptera (beetles) 
49 
13 
Diptera (flies) 
- 
3 
Total number of species 
140 
61 
1 The arachnid groups Pseudoscorpiones, Acari, Opiliones, and Araneae are 
considered subclasses by some workers (see Krantz 1970) and orders by 
others (see Barnes 1980). 
Includes several species whoes ecological status is presently unclear (see text). 
clarification must await additional information on ecology, systematics, 
or both. 
All known cave records within the study area are listed alpha- 
betically by county for each species. Type localities for troglobites are 
indicated in parentheses following the cave name when these localities 
occur in the study area. Quotation marks and a reference (in parentheses) 
to the author who used the name indicate cave localities taken from the 
literature and unknown to either the Virginia Speleological Survey or the 
Tennessee Cave Survey by the name published. Many of the troglobites 
listed are endemic to the study area and, unless indicated otherwise in 
the preliminary discussions or under “Comments,” the caves listed 
