Invertebrate Cave Fauna 
57 
Although to our knowledge this species has not been found outside 
caves to date, its status as a troglobite is uncertain. 
Order Geophilomorpha 
Family Chilenophilidae 
Arctogeophilus umbracticus (McNeill) (AC) 
Virginia. — Scott Co.: Coley Cave No. 2. Shenandoah Co.: Pingleys 
Cave. 
Order Lithobiomorpha 
Family Ethypoliidae 
Bothropolys multidentatus (Newport) (AC) 
Virginia. — Rockbridge Co.: Tolleys Cave. 
Family Lithobiidae 
Nampabius parienus Chamberlin (TX or AC) 
Virginia. — Smyth Co.: Atwells Tunnel Cave. 
Nampabius turbator Crabill (TB?) 
Virginia. — Alleghany Co.: Island Ford and Lowmoor (type locality) 
caves. 
Nampabius sp. 
Virginia. — Montgomery Co.: Erharts Cave. 
Order Scolopendromorpha 
Family Cryptopidae 
Cryptops hortensis Leach (AC) 
Virginia. — Lee Co.: Ruths Cave. 
Crytops hyalinus Say (AC) 
Virginia. — Tazewell Co. Gully Cave. 
Scolopocryptops sexpinosus (Say) (AC) 
Tennessee. — Claiborne Co.: English Cave. 
Theatops posticus (Say) (AC) 
Virginia. — Frederick Co.: Ogdens Cave. 
Class Diplopoda 
Millipeds are among the most common cavernicoles in Virginia and 
east Tennessee and are well represented by a diverse taxonomic 
assemblage consisting of 5 orders, 9 families, 12 genera, and 24 described 
species. Probably about one-half of the species collected from caves are 
undescribed at present. Approximately 25% of the species (including 
both described and undescribed forms) are troglobites. Cavernicolous 
millipeds are usually found in damp to wet areas associated with 
decomposing organic matter (e.g., wood, guano, carcasses) 
Clearly the most significant order with respect to the diversity of 
cavernicolous species and their affinity for the cave environment is the 
Chordeumatida. All of the trogolobitic millipeds in the study area are 
