60 
John R. Holsinger and David C. Culver 
basin northeastward to the Shenandoah Valley (Fig. 22). The genus has 
never been found in caves of the upper Tennessee basin. Troglobitic 
species of Trichopetalum are usually much smaller than those of 
Pseudotremia, and all individuals lack ocelli and pigment. 
Trichopetalum whitei inhabits caves of the Shenandoah Valley and 
is also recorded from caves in adjacent Grant and Pendleton counties, 
W.Va. (see Holsinger et al. 1976). Trichopetalum weyeriensis ranges 
generally south and west of T. whitei and is also recorded from caves in 
Greenbrier, Monroe, Pendleton, and Pocahontas counties, W.Va. (see 
Holsinger et al. 1976). Trichopetalum packardi occurs to the southwest 
of T. weyeriensis and is common in caves of the New River drainage; it 
is also recorded from caves in Greenbrier, Mercer, and Monroe counties, 
W.Va. (Holsinger et al. 1976). 
On the basis of collections made in the early 1960s, N. B. Causey 
(in litt.) concluded that some populations of T. weyeriensis showed 
evidence of intergradation with both T. packardi and T. whitei in 
different parts of West Virginia. Based on these observations, Causey 
(1963) suggested that the three species are subspecies of a single, rather 
widespread species and not three distinct species as she had indicated 
earlier (Causey 1960a). In our judgment, this situation is far from being 
as clear-cut as Causey suggested and cannot be properly resolved until 
all collections from the Virginia-West Virginia cave region (many of 
which have been made since 1963) have been carefully examined and 
analyzed in detail. 
Other chordeumatids recorded from caves in the study area include 
Conotyla venetia (Conotylidae), a possible trogloxene reported from 
one cave and two epigean localities in Alleghany County (see Shear 
1971); and one or more species of Striaria (Striariidae), of which S. 
Columbiana is a possible trogloxene known primarily from epigean 
habitats in northwestern Virginia, adjacent Maryland, and the District 
of Columbia (Chamberlin and Hoffman 1958). One population of 
Striaria from Madisons Saltpetre Cave in Augusta County appears to 
be troglomorphic and may represent an undescribed species, but additional 
study is needed to determine its status vis-a-vis S. columbiana and other 
species in the genus (W. A. Shear, in litt.). 
In the order Julida, Ophyiulus pilosus (Julidae), an introduction 
from Europe and a probable troglophile. is rather widespread and 
occasionally abundant in Virginia caves. It is also recorded from caves 
in Maryland, Ohio, and West Virginia (see Franz and Slifer 1971, 
Holsinger et al. 1976, Hobbs and Flynn 1981). The order Polydesmida 
is poorly represented in study-area caves, and only a few records are 
known. Brachydesmus superus (Polydesmidae), either a trogloxene or 
accidental, is common in Europe and in cultivated areas of the United 
States; it is recorded from single caves in Virginia and West Virginia 
