Invertebrate Cave Fauna 
61 
Fig. 22. Distribution of troglobitic millipeds ( Trichopetalum ) in the study 
area. 
(see Holsinger et al. 1976). Another polydesmid, Scytonotus granulatus, 
probably a trogloxene, is recorded from a single cave in Virginia and 
from caves in Maryland, Ohio, and Pennsylvania (Franz and Slifer 
1971, Holsinger 1976, Hobbs and Flynn 1981); it is widespread over 
most of the eastern half of the United States (Chamberlin and Hoffman 
1958). 
The order Spirostreptida is represented by two species of Cambala 
(Cambalidae). Cambala minor , a troglophile, is widespread in the 
southeastern and east-central United States (Shelley 1979) and has been 
collected from caves in Virginia and eight other states within its range 
(see Loomis 1943, Shear 1969, Holsinger and Peck 1971, Black 1971, 
Craig 1975, Holsinger et al. 1976, Peck and Lewis 1978, Hobbs and 
