Invertebrate Cave Fauna 
87 
Atheta annexa Casey (TP) 
Virginia. — Giles Co.: Giant Caverns. Montgomery Co.: Old Mill 
Cave. Roanoke Co.: Goodwins Cave. Shenandoah Co.: Battlefield 
Crystal Cave. Smyth Co.: Stones Cave No. 2. 
Atheta troglophila Klimaszewski and Peck (TP) 
Virginia. — Lee Co.: Lucy Beatty, Smith, and Young-Fugate caves. 
Aleocharinae (undetermined genus and species) 
Virginia. — Augusta Co.: Glade Cave. 
Brathinus nitidus LeConte (TP or TX) 
Virginia. — Lee Co.: Bowling Cave. Russell Co.: Banners Corner 
Cave. Scott Co.: Coley No. 2 and Flannery caves. 
Quedius { Microsaurus ) erythrogaster Mannerheim (TP) 
Virginia. — Bath Co.: Porters and Roy Lyle caves. Giles Co.: Giant 
(Caverns), Harris and Straleys No. 1 caves. Highland Co.: 
Marshalls Cave. Lee Co.: “Cave No. 1 and No. 3, Pennington 
Gap” (Smetana, 1971:85), and Indian Cave. Rockbridge Co.: 
Doll House Cave. Rockingham Co.: Melrose Cave. Scott Co.: 
Sounding Cave. Shenandoah Co.: Hensleys and Shenandoah 
Wild caves. Smyth Co.: Buchanan Saltpetre Cave. 
Quedius {Microsaurus) mesomelinus (Marsham) (TP or TX) 
Virginia. — Pulaski Co.: Sam Bells Cave. 
Quedius {Microsaurus) spelaeus Horn (TP) 
Virginia.— Bath Co.: Cave Run Pit and Crossroads caves. 
Frederick Co.: Ogdens Cave. Giles Co.: Tawneys Cave. Pulaski 
Co.: Fifty-Foot Hell Cave. Rockingham Co.: Three-D Maze 
Cave. 
Quedius sp. 
Virginia. — Augusta Co.: Grand Caverns and Madisons Saltpetre 
Cave. Lee Co.: Young-Fugate Cave. Page Co.: Luray Caverns. 
Rockingham Co.: Massanutten Caverns. Warren Co.: Skyline 
Caverns. 
Other Staphylinidae 
Nine other genera, based largely on single records and presumably 
including mostly accidental species, are recorded from study-area 
caves as follows: Cratarea, Emplenota, Erichsonius, Homaeotarsus, 
Lathrobium , Lesteva (probably L. pallipes LeConte, a common 
trogloxene), Megalinus, Philonthus, and Trichophya (probably T. 
pilicornis Gyllenhal). 
Order Diptera 
With the possible exception of one species, there are no troglobitic 
dipterans (flies) in the study area. Several species are relatively common 
in caves, however, and sometimes make an important contribution to 
