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John R. Holsinger and David C. Culver 
(2) Some common caves species in the study area, such as the 
spider Meta menardi, the cricket Euhadenoecus fragilis, and the dipterans 
Amoebalaria defessa and Megaselia cavernicola, are apparently well 
adapted in their present roles as troglophiles or (habitual) trogloxenes 
and show no evidence of evolving into troglobites. Other troglophiles, 
however, such as the harvestman Erebomaster acanthina, the spider 
Nesticus carteri, and certain species of the milliped Pseudotremia, show 
good evidence of becoming bona fide troglobites, and, as such, they are 
probably troglobites in statu nascendi. 
ACKNOWLEDGMENTS. — We are extremely grateful to the many 
individuals who have assisted us in various phases of the field work or 
furnished us with useful data from their own collections. In particular 
we thank R. A. Baroody, T. C. Barr, Jr., J. H. Carpenter, J. M. Beck, 
III, L. G. Conrad, J. E. Cooper, G. D. Corbett, J. Cox, G. W. Dickson, 
J. E. Estes, L. M. Ferguson, D. L. Finley, S. W. Hetrick, R. L. 
Hoffman, L. R. Hubricht, T. C. Kane, C. H. Krekeler, P. C. Lucas, 
G. H. Marland, T. G. Marsh, the late D. R. Martin, the late J. P. E. 
Morrison, S. B. Peck, R. D. Powers, Jr., R. M. Norton, J. E. Tichenor, 
V. M. (Tipton) Dalton, R. L. Wallace, and R. E. Whittemore. 
Many systematists aided in the determination of specimens, and 
their expertise and helpful comments are appreciated. We thank: T. C. 
Barr, Jr., beetles; T. E. Bowman, isopods; T. S. Briggs, opilionids; J. H. 
Carpenter, flatworms; the late N. B. Causey, milllipeds; K. A. 
Christiansen, collembolans; D. G. Cook, oligochaetes; R. E. Crabill, Jr., 
centipedes; D. A. Crossley, Jr., mites; W. R. Elliott, mites; L. M. 
Ferguson, diplurans; L. E. Fleming, isopods; B. A. Foote, dipterans; 
R. J. Gagne, dipterans; G. E. Gates, oligochaetes; W. J. Gertsch, 
spiders; C. J. Goodnight, opilionids; A. B. Gurney, crickets; L. J. 
Herman, Jr., beetles; R. Hershler, snails; H. H. Hobbs, Jr., crayfishes; 
R. L. Hoffman, millipeds; P. C. Holt, branchiobdellids; T. H. Hubbell, 
crickets; L. R. Hubricht, snails; L. H. Hyman, flatworms; R. Kenk, 
flatworms; J. J. Lewis, isopods; D. R. Malcolm, pseudoscorpions; S. A. 
Marashall, dipterans; A. C. Mickelbacher, symphylans; the late J. P. E. 
Morrison, snails; W. B. Muchmore, pseudoscorpions; S. B. Peck, 
beetles; G. A. Schultz, isopods; W. A. Shear, millipeds and opilionids; 
D. E. Sonenshine, ticks; H. R. Steeves, III, isopods; G. C. Steyskal, 
dipterans; F. C. Thompson, dipterans; F. G. Thompson, snails; the late 
A. Vandel, isopods; W. C. Welbourn, mites; W. W. Wirth, dipterans; 
D. L. Wray, collembolans; H. C. Yeatman, copepods; and M. Zackarda, 
mites. 
We are indebted to numerous landowners in Virginia and eastern 
Tennessee who so willingly cooperated with us in allowing access to 
