Invertebrate Cave Fauna 
147 
their caves. Many owners went out of their way to assist with the field 
work. We are also grateful to the managements of the privately operated 
commercial caves in Virginia and to officials of Cumberland Gap 
National Historical Park, Natural Tunnel State Park, and the Upper 
Valley Regional Park Authority for giving us free access to work in the 
caves under their jurisdiction. J. R. Jordan, Jr., and W. H. Redman of 
the Regional Heritage Program of the Tennessee Valley Authority 
(TV A) are thanked for their encouragement and support of our research 
in southwestern Virginia and eastern Tennessee. D. L. Miller-Carson, of 
the Old Dominion University Center for Instructional Development, 
and B. A. McBride, of the TVA Division of Natural Resources Services, 
are thanked for their assistance with preparation of the maps. Certain 
parts of this paper have benefited from discussions and/or the exchange 
of information with T. C. Barr, Jr., E. M. Benedict, K. A. Christiansen, 
L. M. Ferguson, F. G. Howarth, T. C. Kane, J. J. Lewis, S. B. Peck, 
T. L. Poulson, J. R. Reddell, F. D. Stone, and many others. 
This study was supported in part by grants from the Research 
Advisory Committee of the National Speleological Society, the National 
Science Foundation (GB-42332), and Old Dominion University (Faculty 
Summer Research Grant in 1979) and by travel funds from the 
Department of Invertebrate Zoology of the Smithsonian Institution 
(summer of 1972) to J. R. Holsinger; by grants from the National 
Science Foundation and Northwestern University to D. C. Culver; and 
by a personal service contract from the Tennessee Valley Authority (TV 
51940A) in 1979 to both of us. This paper is the final contribution of the 
Cave Biogeography of the Central Appalachians Study Group, a former 
research project of the National Speleological Society. 
LITERATURE CITED 
Banks, N. 1895. Notes on the Pseudoscorpionida. J. N.Y. Entomol. Soc. 
3:1-13. 
Barber, Henry S. 1928. Two new cave beetles related to Anophthalmus pusio 
Horn. J. Wash. Acad. Sci. 18:195-196. 
Barnes, Robert D. 1980. Invertebrate Zoology. W. B. Saunders, Philadelphia. 
Barr, Thomas C., Jr. 1959. Speciation and distribution of cavernicolous 
invertebrates in the eastern United States. Yr. Bk. Am. Phil. Soc. 
(1959):229-230. 
Barr, Thomas C., Jr. 1960a. A new genus of cave beetle (Carabidae: Trechini) 
from southwestern Virginia, with a key to the genera of Trechini of North 
America north of Mexico. Coleopt. Bull. 14:65-70. 
Barr, Thomas C., Jr. 1960b. The cavernicolous beetles of the subgenus 
Rhadine, genus Agonum (Coleoptera: Carabidae). Am. Midi. Nat. 64:45-65. 
Barr, Thomas C., Jr. 1961. Caves of Tennessee. Tenn. Div. Geol. Bull. 64, 
Nashville. 
