68 
Thomas C. Barr, Jr. 
tively. A single specimen was found on Fish Camp Prong under moss car- 
peting a low ledge at an elevation of 2200 feet (670 m). On West Prong of 
Little Pigeon River (4000 feet) (1200 m) it occurred under rocks at the 
edge of the stream, with 77 valentinei and Bembidion carolinense Casey. At 
the south base of the Smokies (Swain County, North Carolina) 77 
luculentus was extracted from leaves and wet gravels at 2250 feet (685 m), 
beside a small tributary stream crossing the Deep Creek Trail. A single 
female was collected on Kephart Prong under similar circumstances, at 
an elevation of 3350 feet (1020 m). These collecting data have been given 
in some detail to demonstrate that 77 luculentus is by no means restricted 
to higher elevations, but should be able to bridge the low gaps between 
mountain ranges. A limited amount of gene flow between nominate 
luculentus and the other two subspecies is thus a reasonable proposition, 
although intergrading populations have not yet been discovered. 
Wherever it coexists syntopically with 77 valentinei, T. luculentus is 
always two to three times more abundant. It occasionally is taken 
together with 77 barben and 77 bowhngi , both conspicuously smaller 
species, and rarely with 77 uncifer and 77 novaculosus at higher elevations. 
In the central Smokies the only other large species of Trechus with large 
eyes is 77 nebulosus , an inhabitant of wet, fluffy moss carpets (consequent- 
ly almost never syntopic with 77 luculentus ); in 77 nebulosus the sides of the 
pronotum are simply convergent to the base, and males are readily deter- 
mined by the distinctive shape of the apex of the dorsal copulatory piece 
which often protrudes from the apical orifice of the aedeagus. 
Trechus ( Microtrechus) luculentus umcoi, new subspecies 
Figs. 14, 40, 41 
Trechus ( Microtrechus ) luculentus: Barr 1962:88 (in part). 
Etymology . — From Unicoi Mountains, where the type locality is 
situated. 
Description. — Length 4. 3-4. 5, mean 4.4 mm. Differs from other sub- 
species of T. luculentus in more prominently sinuous pronotum sides, 
which are subparallel in basal 0.12-0.15; more prominent, acute, hind 
angles of pronotum; more extensive longitudinal striation of elytra, with 
all striae usually present, although 5th through 8th progressively 
shallower; and larger aedeagus, 0.95-1.04, mean 1.00 mm long. 
Type series. — Holotype male (AMNH) and 22 paratypes, Stratton 
Meadows, elevation 4900 feet (1494 m), Monroe County, Tennessee, and 
Graham County, North Carolina, 30 August 1964, T. C. Barr. 
