Revision of Trechus 
71 
Smokies). It is known from only two localities: Water Rock Knob, in the 
Plott Balsam Mountains, and Richland Balsam, Haywood County, 
North Carolina, in the Great Balsam Mountains. At both collecting sites 
it was taken deep in the mat of spruce-fir needles piled up against wet, 
vertical rock faces. It is easily distinguished from the other species of 
Trechus with which it is sympatric (77 aduncus, T. balsamensis, T. barben, T. 
subtilis, T. vandykei) by size alone. 
Trechus ( Microtrechus) tuckaleechee Barr 
Fig. 16 
Barr 1962:86, Fig. 22. Type locality, Tuckaleechee Caverns, Blount 
County, Tennessee; type deposited in USNM. 
This species is known only from Tuckaleechee Caverns near Town- 
send, Tennessee, at the north side of the Great Smoky Mountains. It is 
distinguished from other members of the nebulosus group by small eyes, 
sinuate sides of the pronotum, hind angles sharp and nearly right, elytra 
with five clearly impressed striae and short apical groove, and the apex of 
the aedeagus broad, finely reflexed at the very tip. Morphologically it ap- 
pears to be closest to T. tennesseensis and T. nantahalae. 
Trechus ( Microtrechus ) tennesseensis tennesseensis Barr 
Barr 1962:87, Fig. 23. Type locality, Berry Cave, Roane County, Ten- 
nessee; type deposited in USNM. 
This taxon is at present known from a single locality in the floor of 
the Appalachian valley approximately 50 miles (80 km) west of the 
Great Smoky Mountains. Berry Cave is located a quarter of a mile (0.4 
km) west of the Tennessee River (mile 578.4) near Wright Bend, 0.15 
mile (0.24 km) west of Tennessee Route 72, at an elevation of 840 feet 
(255 m). Trechus t. tennesseensis resembles T. tuckaleechee , but the hind 
angles of the pronotum are sharp and acute, and the aedeagus is much 
larger. Neither T. tuckaleechee nor T. tennesseensis exhibit any regressive 
modifications that would restrict them to cave habitats. Presumably 
they are both relics of species that were more widely distributed during 
glacial maxima. 
Trechus ( Microtrechus ) tennesseensis tauricus Barr 
Barr 1962: 87. Type locality, Bull Cave, Blount County, Tennessee; type 
deposited in USNM. 
This subspecies, known only from the sinkhole at the entrance to Bull 
