New Cave Planarian 
93 
Fig. 2. Phagocata carolinensis, semidiagrammatic view of copulatory apparatus in sagittal 
sectiop. ac, common atrium; am , male atrium; b, copulatory bursa; bd , bursal duct; bp, penis 
bulb; gp, gonopore; la, lamellae; m, mouth; ode, common oviduct; ph, pharyngeal pouch; pp, 
penis papilla; va, vagina; vd, vas deferens. 
muscles. In the region of the pharynx, the vasa deferentia expand to form 
the tortuous false seminal vesicles or spermiductal vesicles, filled with 
sperm, which proceed posteriorly toward the copulatory complex. 
The copulatory apparatus (Fig. 2) is located immediately behind the 
pharyngeal pouch {ph). The genital aperture or gonopore {gp) is situated 
about midway between the mouth {m) and the posterior end of the body. 
It leads through a short narrow duct into the common atrium {ac) which 
opens anteriorly into the male atrium {am) and posteriorly into the vagina 
{va) of the bursal duct. Both the male and common atria are lined with a 
nucleate and ciliated cuboidal epithelium. 
The penis consists of a rather voluminous bulb {bp), the musculature of 
which is rather feeble and developed mainly near its periphery, and a 
short, bluntly conical papilla {pp) extending into the male atrium. The 
configuration of the penial lumen is quite unique. Three to five transverse 
lamellae {la) originate from the dorsal and ventral walls, alternatingly 
projecting downward and upward. Each lamella is attached to the lateral 
walls of the penis, as can be seen in horizontal sections through the organ. 
