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Roman Kenk 
Fig. 3. Photomicrographs of copulatory organs in sagittal section. A, Phagocata carolinensis, 
X60; B, Phagocata holleri, X127. 
This arrangement would permit the passage of sperm from the bulb to the 
tip of the penis papilla only in a serpentine fashion. It is possible, 
however, that the configuration of the penis lumen as it appears in the 
slides obtains only when the penis is retracted. When the papilla is ex- 
tended during copulation the passage may well straighten out. Both the 
lamellae and the greater part of the outer surface of the papilla are 
covered by a thin infranucleate epithelium with two underlying muscle 
layers, a circular and a longitudinal one. There is no differentiation of the 
penial lumen into a seminal vesicle and ejaculatory duct. At most, a small 
anterior portion of the lumen may differ from the remaining part by hav- 
ing a ciliated and nucleate epithelium. The vasa deferentia (vd) enter the 
penis bulb ventrolaterally, turn dorsomedially, and unite at their opening 
into the anterior end of the penial lumen. 
The copulatory bursa (b) adjoins the wall of the pharyngeal pouch 
(ph). Its outlet, the bursal stalk or duct (bd) proceeds posteriorly for a 
short distance as a narrow duct situated somewhat to the left of the 
midline and lined with a nucleate, ciliated epithelium. It then expands 
into a large, irregularly lobed sac, the vagina (va), the greater part of 
